Although chiefly arboreal, many of the iguanas take readily to the
water; and there is at least one species, _Amblyrhynchus cristatus_,
which leads for the most part an aquatic life. These marine lizards
occur only in the Galapagos Islands, where they are never seen more than
20 yds. inland, while they may often be observed in companies several
hundreds of yards from the shore, swimming with great facility by means
of their flattened tails. Their feet are all more or less webbed, but in
swimming they are said to keep these organs motionless by their sides.
Their food consists of marine vegetation, to obtain which they dive
beneath the water, where they are able to remain, without coming to the
surface to breathe, for a very considerable time. Though they are thus
the most aquatic of lizards, Darwin, who studied their habits during his
visit to those islands, states that when frightened they will not enter
the water. Driven along a narrow ledge of rock to the edge of the sea,
they preferred capture to escape by swimming, while if thrown into the
water they immediately returned to the point from which they started. A
land species belonging to the allied genus _Conolophus_ also occurs in
the Galapagos, which differs from most of its kind in forming burrows in
the ground.
IGUANODON, a large extinct herbivorous land reptile from the Wealden
formation of western Europe, almost completely known by numerous
skeletons from Bernissart, near Mons, Belgium. It is a typical
representative of the ornithopodous (Gr. for bird-footed) Dinosauria.
The head is large and laterally compressed with a blunt snout, nearly
terminal nostrils and relatively small eyes. The sides of the jaws are
provided with a close series of grinding teeth, which are often worn
down to stumps; the front of the jaws forms a toothless beak, which
would be encased originally in a horny sheath. When unworn the teeth are
spatulate and crimped or serrated round the edge, closely resembling
those of the existing Central American lizard, _Iguana_--hence the name
_Iguanodon_ (Gr. Iguana-tooth) proposed by Mantell, the discoverer of
this reptile, in 1825. The bodies of the vertebrae are solid; and they
are convexo-concave (i.e. _opisthocoelous_) in the neck and anterior
part of the back, where there must have been much freedom of motion. The
hindquarters are comparatively large and heavy, while the tail is long,
deep and more or less laterally compressed, evidently adapted for
swimming. The small and mobile fore-limbs bear four complete fingers,
with the thumb reduced to a bony spur. The pelvis and hind-limbs much
resemble those of a running bird, such as those of an emu or the extinct
moa; but the basal bones (metatarsals) of the three-toed foot remain
separate throughout life, thus differing from those of the running
birds, which are firmly fused together even in the young adult. No
external armour has been found. The reptile doubtless frequented
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