Selachians.--No parapineal organ is present. The pineal body (except
in _Torpedo_ where it is absent) is in the form of a long slender tube
ending in front in a dilated bulb lying near the front end of the
brain in close contact with, or enclosed in, a definite foramen in the
cranial roof.
Holocephali and Crossopterygii.--Here also the pineal body is long and
tubular: at its origin it passes dorsalwards or slightly backwards
behind the large dorsal sac.
Actinopterygian Ganoids resemble Selachians on the whole. In _Amia_ a
parapineal organ is present, and it is said to lie towards the left
side and to be connected by a thick nerve with the _left_ habenular
ganglion (cf. _Petromyzon_, article CYCLOSTOMATA). This is adduced to
support the view that the pineal and parapineal bodies represent
originally paired structures.
Teleostei.--A parapineal rudiment appears in the embryo of some forms,
but in the adult only the pineal organ is known to exist. This is
usually short and club-shaped, its terminal part with much folded wall
and glandular in character. In a few cases a parietal foramen occurs
(_Callichthys_, _Loricaria_, &c.).
Dipneusti.--The pineal organ is short and simple. No parapineal organ
is developed.
The dorsal sac is formed by that part of the roof of the
thalamencephalon lying between the habenular commissure and the region
of the velum. In some cases a longitudinal groove is present in which
the pineal organ lies (Dipneusti). In the Crossopterygians the dorsal
sac is particularly large and was formerly mistaken for the pineal
organ.
The _velum transversum_ is a transverse, inwardly-projecting fold of the
roof of the primitive fore-brain in front of the dorsal sac. To those
morphologists who regard the hemisphere region or telencephalon as a
primitively unpaired structure the velum is an important landmark
indicating the posterior limit of the telencephalon. Those who hold the
view taken in this article that the hemispheres are to be regarded as
paired outpushings of the side wall of the primitive fore-brain
attribute less morphological importance to the velum. Physiologically
the velum is frequently important from the plexus of blood-vessels which
passes with it into the III. ventricle.
In _Petromyzon_ and _Chimaera_ the velum is not developed. In Dipnoans
there are present in its place _paired_ transverse folds which are
probably merely extensions backwards of the lateral plexuses.
Public-domain text, read in full here on John Shaqi.
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