The _eyes_ are usually of large size. The lens is large and spherical
and in the case of most Teleostomes accommodation for distant vision is
effected by the lens being pulled bodily nearer the retina. This
movement is brought about by the contraction of smooth muscle fibres
contained in the _processus falciformis_, a projection from the choroid
which terminates in contact with the lens in a swelling, the _campanula
Halleri_. In _Amia_ and in Teleosts a network of capillaries forming the
so-called choroid gland surrounds the optic nerve just outside the
retina. As a rule the eyes of fishes have a silvery, shining appearance
due to the deposition of shining flakes of guanin in the outer layer of
the choroid (_Argentea_) or, in the case of Selachians, in the inner
layers (_tapetum_). Fishes which inhabit dark recesses, e.g. of caves or
of the deep sea, show an enlargement, or, more frequently, a reduction,
of the eyes. Certain deep-sea Teleosts possess remarkable telescopic
eyes with a curious asymmetrical development of the retina.[49]
The otocyst or auditory organ agrees in its main features with that of
other vertebrates. In Selachians the otocyst remains in the adult open
to the exterior by the _ductus endolymphaticus_. In _Squatina_[50] this
is unusually wide and correlated; with this the calcareous otoconia are
replaced by sand-grains from the exterior. In Dipnoans (_Lepidosiren_
and _Protopterus_) curious outgrowths arise from the ductus
endolymphaticus and come to overlie the roof of the fourth ventricle,
recalling the somewhat similar condition met with in certain Amphibians.
In various Teleosts the swimbladder enters into intimate relations
with the otocyst. In the simplest condition these relations consist in
the prolongation forwards of the swimbladder as a blindly ending tube
on either side, the blind end coming into direct contact either with
the wall of the otocyst itself or with the fluid surrounding it
(perilymph) through a gap in the rigid periotic capsule. A wave of
compression causing a slight inward movement of the swimbladder wall
will bring about a greatly magnified movement of that part of the wall
which is not in relation with the external medium, viz. the part in
relation with the interior of the auditory capsule. In this way the
perception of delicate sound waves may be rendered much more perfect.
In the Ostariophysi (Sagemehl), including the _Cyprinidae_, the
_Siluridae_, the _Characinidae_ and the _Gymnotidae_, a
physiologically similar connexion between swimbladder and otocyst is
brought about by the intervention of a chain of auditory ossicles
(Weberian ossicles) formed by modification of the anterior
vertebrae.[51]
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