In the Carboniferous period, the Ostracophori and Arthrodira have
disappeared, the Crossopterygii and Dipneusti are still abundant, and
the Selachians (_Pleuracanthus_, Acanthodians, truesharks) and
Chondrostean ganoids (_Palaeoniscidae_ and _Platysomidae_) are
predominant. In the Upper Permian the Holostean ganoids
(_Acanthophorus_) make their appearance, and the group becomes dominant
in the Jurassic and the Lower Cretaceous. In the Trias, the
Crossopterygii and Dipneusti dwindle in variety and the _Ceratodontidae_
appear; the Chondrostean and Holostean ganoids are about equally
represented, and are supplemented in the Jurassic by the first,
annectant representatives of the Teleostei (_Pholidophoridae_,
_Leptolepidae_). In the latter period, the Holostean ganoids are
predominant, and with them we find numerous Cestraciont sharks, some
primitive skates (_Squatinidae_ and _Rhinobatidae_), Chimaerids and
numerous Coelacanthid crossopterygians.
The fish-fauna of the Lower Cretaceous is similar to that of the
Jurassic, whilst that of the Chalk and other Upper Cretaceous formations
is quite modern in aspect, with only a slight admixture of Coelacanthid
crossopterygians and Holostean ganoids, the Teleosteans being abundantly
represented by _Elopidae_, _Albulidae_, _Halosauridae_, _Scopelidae_ and
_Berycidae_, many being close allies of the present inhabitants of the
deep sea. At this period the spiny-rayed Teleosteans, dominant in the
seas of the present day, made their first appearance.
With the Eocene, the fish-fauna has assumed the essential character
which it now bears. A few Pycnodonts survive as the last representatives
of typically Mesozoic ganoids, whilst in the marine deposits of Monte
Bolca (Upper Eocene) the principal families of living marine fishes are
represented by genera identical with or more or less closely allied to
those still existing; it is highly remarkable that forms so highly
specialized as the sucking-fish or remoras, the flat-fish
(_Pleuronectidae_), the Pediculati, the Plectognaths, &c., were in
existence, whilst in the freshwater deposits of North America
_Osteoglossidae_ and _Cichlidae_ were already represented. Very little
is known of the freshwater fishes of the early Tertiaries. What has been
preserved of them from the Oligocene and Miocene shows that they
differed very slightly from their modern representatives. We may
conclude that from early Tertiary times fishes were practically as they
are at present. The great hiatus in our knowledge lies in the period
between the Cretaceous and the Eocene.
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