A division of the world into regions according to the distribution of
the shore-fishes is a much more difficult task than that of tracing
continental areas. It is possible perhaps to distinguish four great
divisions: the Arctic region, the Atlantic region, the Indo-Pacific
region and the Antarctic region. The second and third may be again
subdivided into three zones: Northern, Tropical and Southern. This
appears to be a more satisfactory arrangement than that which has been
proposed into three zones primarily, each again subdivided according to
the different oceans. Perhaps a better division is that adopted by D. S.
Jordan, who arranges the littoral fishes according to coast lines; we
then have an East Atlantic area, a West Atlantic, an East Pacific and a
West Pacific, the latter including the coasts of the Indian Ocean. The
tropical zone, whatever be the ocean, is that in which fishes flourish
in greatest abundance and where, especially about coral-reefs, they show
the greatest variety of bizarre forms and the most gorgeous coloration.
The fish-fauna of the Indo-Pacific is much richer than that of the
Atlantic, both as regards genera and species.
As regards the Arctic and Antarctic regions, the continuity or
circumpolar distribution of the shore fishes is well established. The
former is chiefly characterized by its Cottids, Cyclopterids, Zoarcids
and Gadids, the latter by its Nototheniids. The theory of bipolarity
receives no support from the study of the fishes.
Pelagic fishes, among which we find the largest Selachians and
Teleosteans, are far less limited in their distribution, which, for many
species, is nearly world-wide. Some are dependent upon currents, but the
great majority being rapid swimmers able to continue their course for
weeks, apparently without the necessity of rest (many sharks, scombrids,
sword-fishes), pass from one ocean into the other. Most numerous between
the tropics, many of these fishes occasionally wander far north and
south of their habitual range, and there are few genera that are at all
limited in their distribution.
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