More distinct is the Papuan genus _Dorcopsis_, as typified by _D.
muelleri_, although it is to some extent connected with _Macropus_ by
_D. macleyi_. The muzzle is naked, the fur on the nape of the neck
directed more or less completely forward, and the hind-limbs are less
disproportionately elongated. Perhaps, however, the most distinctive
feature of the genus is the great fore-and-aft length of the
penultimate premolar in both jaws. Other species are _D.
rufolateralis_ and _D. aurantiacus_. In the tree-kangaroos, which
include the Papuan _Dendrolagus inustus_, _D. ursinus_, _D. dorianus_,
_D. benetianus_ and _D. maximus_, and the North Queensland _D.
lumholtzi_, the reduction in the length of the hind-limbs is carried
to a still further degree, so that the proportions of the fore and
hind limbs are almost normal. The genus agrees with _Dorcopsis_ in the
direction of the hair on the neck, but the muzzle is only partially
hairy, and the elongation of the penultimate premolar is less. These
kangaroos are largely arboreal in their habits, but they descend to
the ground to feed. Lastly, we have the banded wallaby, _Lagostrophus
fasciatus_, of Western Australia, a small species characterized by its
naked muzzle, the presence of long bristles on the hind-feet which
conceal the claws, and also of dark transverse bands on the lower part
of the back. The skull has a remarkably narrow and pointed muzzle and
much inflated auditory bullae; while the two halves of the lower jaw
are firmly welded together at their junction, thus effectually
preventing the scissor-like action of the lower incisors distinctive
of _Macropus_ and its immediate allies. As regards the teeth, canines
are wanting, and the penultimate upper premolar is short, from before
backwards, with a distinct ledge on the inner side.
In the rat-kangaroos, or kangaroo-rats, as they are called in
Australia, constituting the sub-family _Potoroinae_, the first upper
incisor is narrow, curved, and much exceeds the others in length; the
upper canines are persistent, flattened, blunt and slightly curved,
and the first two premolars of both jaws have large, simple,
compressed crowns, with a nearly straight or slightly concave free
cutting-edge, and both outer and inner surfaces usually marked by a
series of parallel, vertical grooves and ridges. Molars with quadrate
crowns and a blunt conical cusp at each corner, the last notably
smaller than the rest, sometimes rudimentary or absent. Forefeet
narrow; the three middle toes considerably exceeding the first and
fifth in length and their claws long, compressed and but slightly
curved. Hind-feet as in _Macropus_. Tail long, and sometimes partially
prehensile when it is used for carrying bundles of grass with which
these animals build their nests. The group is confined to Australia
and Tasmania, and all the species are relatively small.
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