here by hypothesis an elongated neck; it is conceivable that out of
large herds the few survivors of a drought might be exclusively such as
possessed this advantage to some extent. These would probably transmit
to a large majority of their descendants the tendency to vary in a
given direction which they had themselves all more or less exhibited.
Their progeny, moreover, would be placed in exceptionally favourable
circumstances by the very fact that in the previous drought so many of
the same species had been starved to death, who would otherwise have
furnished their chief competitors in the struggle for existence. It
is still objected that upon this supposition many other animals ought
to have acquired giraffe-like necks. But such an expectation is far
from being warranted by the principles of Natural Selection. Since
all variations are potentially useful, but only those are preserved
which suit the surrounding conditions among which they are exhibited,
the calculation of chances will itself plead for the probability that
a variety of variations will be preserved, rather than the same many
times over. Other species competing with the giraffe for food would
be little likely to gain an advantage over it by a slight increase in
length of neck, though by other variations they might achieve a decided
superiority. It is obvious, also, that the advantage assigned to the
elongated neck would belong to many other possible variations, such as
a lengthened proboscis, far-reaching arms, the climbing powers of the
snake or the monkey, the flight of the bird or the insect; all of which
may be due to Natural Selection and the subsidiary principles which the
theory of Development embraces.
The calling in of subsidiary principles may be thought to spoil the
boasted simplicity of the theory. But such an opinion is hypercritical.
One might truthfully say of a great patriot that all he did was in
obedience to the simple law of duty, without implying that he was
exempt from the law of association of ideas, or independent of the
mechanical, chemical, and vital laws which regulate many of the
functions of all human beings alike.
FOOTNOTES.
[1] These papers were read at various dates between February 1869 and
January 1871, before the Torquay Natural History Society.
[2] Read before the Devonshire Association, July 1870.
[3] Horace, Odes, I. xvi. 13:--
Fertur Prometheus addere principi
Limo coactus particulam undique
Desectam, et insani leonis
Vim stomacho apposuisse nostro.
[4] Lyell, ‘Principles of Geology,’ vol. ii. p. 324. Tenth Edition.
[5] Darwin, ‘Animals and Plants under Domestication,’ vol. ii. p. 251.
[6] 1,073,741,824 grains.
[7] ‘On the Origin of Species,’ p. 74. Fourth Edition.
[8] Darwin, ‘Animals and Plants under Domestication,’ vol. ii. p. 379.
[9] 37 dwts. 7 grs., or 895 grs., between seven and eight times the
size of the wild fruit. See ‘Animals and Plants under Domestication,’
vol. i. p. 356.
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