The question at issue between the Lamarckians, supported by Mr. Spencer and a growing band of those who have risen in rebellion against the Charles-Darwinian system on the one hand, and Messrs. Darwin and Wallace with the greater number of our more prominent biologists on the other, involves the very existence of evolution as a workable theory. For it is plain that what Nature can be supposed able to do by way of choice must depend on the supply of the variations from which she is supposed to choose. She cannot take what is not offered to her; and so again she cannot be supposed able to accumulate unless what is gained in one direction in one generation, or series of generations, is little likely to be lost in those that presently succeed. Now variations ascribed mainly to use and disuse can be supposed capable of being accumulated, for use and disuse are fairly constant for long periods among the individuals of the same species, and often over large areas; moreover, conditions of existence involving changes of habit, and thus of organisation, come for the most part gradually; so that time is given during which the organism can endeavour to adapt itself in the requisite respects, instead of being shocked out of existence by too sudden change. Variations, on the other hand, that are ascribed to mere chance cannot be supposed as likely to be accumulated, for chance is notoriously inconstant, and would not purvey the variations in sufficiently unbroken succession, or in a sufficient number of individuals, modified similarly in all the necessary correlations at the same time and place to admit of their being accumulated. It is vital therefore to the theory of evolution, as was early pointed out by the late Professor Fleeming Jenkin and by Mr. Herbert Spencer, that variations should be supposed to have a definite and persistent principle underlying them, which shall tend to engender similar and simultaneous modification, however small, in the vast majority of individuals composing any species. The existence of such a principle and its permanence is the only thing that can be supposed capable of acting as rudder and compass to the accumulation of variations, and of making it hold steadily on one course for each species, till eventually many havens, far remote from one another, are safely reached.