"In such a case we have only to assume that the climate which is
unfavourable, and the nutriment which is insufficient for horses, affect
not only the animal as a whole but also its germ-cells. This would
result in the diminution in size of the germ-cells, the effects upon the
offspring being still further intensified by the insufficient nourishment
supplied during growth. But such results would not depend upon the
transmission by the germ-cells of certain peculiarities due to the
unfavourable climate, which only appear in the full-grown horse."
But Professor Weismann does not like such cases, and admits that he
cannot explain the facts in connection with the climatic varieties of
certain butterflies, except "by supposing the passive acquisition of
characters produced by the direct influence of climate."
Nevertheless in his next paragraph but one he calls such cases
"doubtful," and proposes that for the moment they should be left aside.
He accordingly leaves them, but I have not yet found what other moment he
considered auspicious for returning to them. He tells us that "new
experiments will be necessary, and that he has himself already begun to
undertake them." Perhaps he will give us the results of these
experiments in some future book--for that they will prove satisfactory to
him can hardly, I think, be doubted. He writes:--
"Leaving on one side, for the moment, these doubtful and insufficiently
investigated cases, we may still maintain that the assumption that
changes induced by external conditions in the organism as a whole are
communicated to the germ-cells after the manner indicated in Darwin's
hypothesis of pangenesis, is wholly unnecessary for the explanation of
these phenomena. Still we cannot exclude the possibility of such a
transmission occasionally occurring, for even if the greater part of the
effects must be attributable to natural selection, there might be a
smaller part in certain cases which depends on this exceptional factor."
Public-domain text, read in full here on John Shaqi.
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