the germ-cells that they hand down its scars to offspring, how much more
shall not anxieties that have directed action of all kinds from birth
till death, not in one generation only but in a longer series of
generations than the mind can realise to itself, modify, and indeed
control, the organisation of every species?
I see Professor S. H. Vines, in the article on Weismann's theory referred
to in my preceding article, says Mr. Darwin "held that it was not the
sudden variations due to altered external conditions which become
permanent, but those slowly produced by what he termed 'the accumulative
action of changed conditions of life.'" Nothing can be more soundly
Lamarckian, and nothing should more conclusively show that, whatever else
Mr. Darwin was, he was not a Charles-Darwinian; but what evidence other
than inferential can from the nature of the case be adduced in support of
this, as I believe, perfectly correct judgment? None know better than
they who clamour for direct evidence that their master was right in
taking the position assigned to him by Professor Vines, that they cannot
reasonably look for it. With us, as with themselves, modification
proceeds very gradually, and it violates our principles as much as their
own to expect visible permanent progress, in any single generation, or
indeed in any number of generations of wild species which we have yet had
time to observe. Occasionally we can find such cases, as in that of
_Branchipus stagnalis_, quoted by Mr. Wallace, or in that of the New
Zealand Kea whose skin, I was assured by the late Sir Julius von Haast,
has already been modified as a consequence of its change of food. Here
we can show that in even a few generations structure is modified under
changed conditions of existence, but as we believe these cases to occur
comparatively rarely, so it is still more rarely that they occur when and
where we can watch them. Nature is eminently conservative, and fixity of
type, even under considerable change of conditions, is surely more
important for the well-being of any species than an over-ready power of
adaptation to, it may be, passing changes. There could be no steady
progress if each generation were not mainly bound by the traditions of
those that have gone before it. It is evolution and not incessant
revolution that both parties are upholding; and this being so, rapid
visible modification must be the exception, not the rule. I have quoted
direct evidence adduced by competent observers, which is, I believe,
sufficient to establish the fact that offspring can be and is sometimes
modified by the acquired habits of a progenitor. I will now proceed to
the still more, as it appears to me, cogent proof afforded by general
considerations.
What, let me ask, are the principal phenomena of heredity? There must be
physical continuity between parent, or parents, and offspring, so that
the offspring is, as Erasmus Darwin well said, a kind of elongation of
the life of the parent.
Public-domain text, read in full here on John Shaqi.
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