To this cause of minor breaks in the succession of organic forms--a cause on which we have dwelt because it has not been taken into account--we must add sundry others. Besides these periodically-recurring changes of climate, there are the irregular ones produced by redistributions of land and sea; and these, sometimes less, sometimes greater, in degree, than the rhythmical changes, must, like them, cause in each region emigrations and immigrations of species; and consequent breaks, small or large as the case may be, in the paleontological series. Other and more special geological changes must produce other and more local blanks in the succession. By some inland elevation the natural drainage of a continent is modified; and instead of the sediment previously brought down to the sea by it, a great river brings down sediment unfavourable to various plants and animals living in its delta: whereupon these disappear from the locality, perhaps to re-appear in a changed form after a long epoch. Upheavals or subsidences of shores or sea-bottoms, involving deviations of marine currents, remove the habitats of many species to which such currents are salutary or injurious; and further, this redistribution of currents alters the places of sedimentary deposits, and thus stops the burying of organic remains in some localities, while commencing it in others. Had we space, many more such causes of blanks in our paleontological records might be added. But it is needless here to enumerate them. They are admirably explained and illustrated in Sir Charles Lyell's _Principles of Geology_. Now, if these minor changes of the Earth's surface produce minor breaks in the series of fossilized remains; must not great changes produce great breaks? If a local upheaval or subsidence causes throughout its small area the absence of some links in the chain of fossil forms; does it not follow that an upheaval or subsidence extending over a large part of the Earth's surface, must cause the absence of a great number of such links throughout a very wide area?