Essays Upon Heredity and Kindred Biological Problems: Authorised TranslationWeismann, August
Science
Essays Upon Heredity and Kindred Biological Problems: Authorised Translation
Weismann, August
Evolution (Biology); Genetics; Reproduction
I must here break off in the midst of these considerations, rather than
conclude them, for much still remains to be said. I hope, nevertheless,
that I have thrown new light upon some important points, and I now
propose to conclude with the following short abstract of the results of
my enquiry.
I. Natural death occurs only among multicellular beings; it is not
found among unicellular organisms. The process of encystment in the
latter is in no way comparable with death.
II. Natural death first appears among the lowest Heteroplastid Metazoa,
in the limitation of all the cells collectively to one generation, and
of the somatic or body-cells proper to a restricted period: the somatic
cells afterwards in the higher Metazoa came to last several and even
many generations, and life was lengthened to a corresponding degree.
III. This limitation went hand in hand with a differentiation of the
cells of the organism into reproductive and somatic cells, in
accordance with the principle of division of labour. This
differentiation took place by the operation of natural selection.
IV. The fundamental biogenetic law applies only to multicellular
beings; it does not apply to unicellular forms of life. This depends on
the one hand upon the mode of reproduction by fission which obtains
among the Monoplastides (unicellular organisms), and on the other upon
the necessity, induced by sexual reproduction, for the maintenance of a
unicellular stage in the development of the Polyplastides
(multicellular organisms).
V. Death itself, and the longer or shorter duration of life, both
depend entirely on adaptation. Death is not an essential attribute of
living matter; it is neither necessarily associated with reproduction,
nor a necessary consequence of it.
* * * * *
In conclusion, I should wish to call attention to an idea which is
rather implied than expressed in this essay:—it is, that reproduction
did not first make its appearance coincidently with death. Reproduction
is in truth an essential attribute of living matter, just as is the
growth which gives rise to it. It is as impossible to imagine life
enduring without reproduction as it would be to conceive life lasting
without the capacity for absorption of food and without the power of
metabolism. Life is continuous and not periodically interrupted: ever
since its first appearance upon the earth, in the lowest organisms, it
has continued without break; the forms in which it is manifested have
alone undergone change. Every individual alive to-day—even the very
highest—is to be derived in an unbroken line from the first and lowest
forms.
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Footnotes for Chapter III.
Footnote 59:
‘Ueber den Ursprung des Todes,’ Hamburg and Leipzig, 1883.
Footnote 60:
Public-domain text, read in full here on John Shaqi.
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