Essays Upon Heredity and Kindred Biological Problems: Authorised TranslationWeismann, August
Science
Essays Upon Heredity and Kindred Biological Problems: Authorised Translation
Weismann, August
Evolution (Biology); Genetics; Reproduction
We do not know the number of generations over which the specific
hereditary tendencies of the first generation can make themselves felt.
Many facts seem to indicate however that the number is large, and it is
at all events greater than six. When we remember that, in the tenth
generation, a single germ contains 1024 different germ-plasms, with
their inherent hereditary tendencies, it is quite clear that continued
sexual reproduction can never lead to the re-appearance of exactly the
same combination, but that new ones must always arise.
New combinations are all the more probable because the different
idioplasms composing the germ-plasm in the germ-cells of any individual
are present in different degrees of intensity at different times of its
life; in other words, the intensity of the component idioplasms is a
function of time. This conclusion follows from the fact that children
of the same parents are never exactly identical. In one child the
characters of the father may predominate, in another those of the
mother, in another again those of either grand-parent or
great-grand-parent.
We are thus led to the conclusion that even in a few sexually produced
generations a large number of well-marked individuals must arise: and
this would even be true of generations springing from our hypothetical
species, assumed to be without ancestors, and characterised by few
individual differences. But of course organisms which reproduce
themselves sexually are never without ancestors, and if these latter
were also propagated by the sexual method, it follows that each
generation of every sexual species is in the stage which we have
previously assumed for the tenth or some much later generation of the
hypothetical species. In other words, each individual contains a
maximum of hereditary tendencies and an infinite variety of possible
individual characters (see Appendix VI, p. 326).
In this manner we can explain the origin of hereditary individual
variability as it is known in man and the higher animals, and as it is
required for the theory which explains the transformation of species by
means of natural selection.
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