Essays Upon Heredity and Kindred Biological Problems: Authorised TranslationWeismann, August
Science
Essays Upon Heredity and Kindred Biological Problems: Authorised Translation
Weismann, August
Evolution (Biology); Genetics; Reproduction
There is again the frequently-quoted instance of the young pointer,
‘which, untrained, and without any example which might have been
imitated, pointed at a lizard in a subtropical jungle, just as many of
its forefathers had pointed at partridges on the plain of St. Denis,’
and which, without knowing the effect of a shot, sprang forward
barking, at the first discharge, to bring in the game. This conduct
must not be attributed to the inheritance of any mental picture, such
as the effect of a shot, but to the inheritance of a certain reflex
mechanism. The young pointer does not spring forward at the shot
because he has inherited from his forefathers a certain association of
ideas,—shot and game,—but because he has inherited a reflex mechanism,
which impels him to start forward on hearing a report. We cannot yet
determine without more experiments how such an impulse due to
perception (‘Wahrnehmungstrieb,’ Schneider) has arisen; but, in my
opinion, it is almost inconceivable that artificial breeding has had
nothing to do with it; and that we are here concerned—not with the
inheritance of the effects of training—but with some predisposition on
the part of the germ, which has been increased by artificial selection.
The necessity for extreme caution in appealing to the supposed
hereditary effects of use, is well shown in the case of those numerous
instincts, which only come into play once in a lifetime, and which do
not therefore admit of improvement by practice. The queen-bee takes her
nuptial flight only once, and yet how many and complex are the
instincts and the reflex mechanisms which come into play on that
occasion. Again, in many insects the deposition of eggs occurs but once
in a life-time, and yet such insects always fulfil the necessary
conditions with unfailing accuracy, either simply dropping the eggs
into water, or carefully fixing them on the surface of the earth
beneath some stone, or laying them on a particular part of a certain
species of plant; and in all these cases the most complicated actions
are performed. It is indeed astonishing to watch one of the _Cynipidae_
(_Rhodites rosae_) depositing her eggs in the tissue of a young bud.
She first carefully examines the bud on all sides, and feels it with
her legs and antennae. Then she slowly inserts her long ovipositor
between the closely-rolled leaves of the bud, but if it does not reach
exactly the right spot, she will withdraw and re-insert it many times,
until at length, when the proper place has been found, she will slowly
bore deep into the very centre of the bud, so that the egg will reach
the exact spot, where the necessary conditions for its development
alone exist.
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