Everyday Objects; Or, Picturesque Aspects of Natural History.Adams, W. H. Davenport (William Henry Davenport)
History
Everyday Objects; Or, Picturesque Aspects of Natural History.
Adams, W. H. Davenport (William Henry Davenport)
Natural history
The _tetrapetalous_ (or four-petalled) corolla is usually arranged
like a cross, and is much more frequent than the _dipetalous_ or
_tripetalous_; for examples, we need only refer to the large and
important family of the Cruciferæ.
The number five (_pentapetalous_) is still more common; but we meet
with it in other organs besides petals, and it seems particularly
characteristic of the vegetable kingdom.
Thus, all the Umbelliferæ have five sepals, five petals, and five
stamens; in all the Crassulaceæ, the number five applies, not only to
the sepals and the petals, as well as to the stamens, but also to the
carpels which compose the ovary.
Corollas with six, eight, nine, ten, or twelve petals are relatively
rarer; and when the petals become so numerous that we cannot count
them, we have to deal with transformations of stamens into petals,
with those monstrosities of cultivation which we call double flowers,
_flores pleni_, where all the male organs have disappeared,--flowers
wholly unfit for fructification.
The petals of the corolla are not always free. Like those of the
calyx, they may be attached to one another by their edges, but this
union invariably takes place, as in the former, from bottom to top.
Therefore, we never see any petals united at the top, and disengaged
at the bottom (see Fig. 47, _a_.) But the reader must take careful
note that this invariable characteristic is not peculiar only to the
floral envelopes; it is not met with in the stamens, for these may
be united, either by their anthers, as in the whole family of the
Synantheræ, or by their filaments, as in the Leguminosæ. And what
we have said of the stamens applies also to the parts constituting
the pistil. This radical difference between the perianth and the
true reproductive organs ought, from the beginning, to have fixed
the attention of botanists on the centripetal and centrifugal
metamorphosis of the leaf, which we have spoken of in "The Circle of
the Year."
[Illustration: FIG. 47, _a_.--Natural junction of Petal.]
[Illustration: FIG. 47, _b_.--Unnatural (and impossible) junction.]
* * * * *
In many plants we are permitted to follow step by step, as it were,
the union of the petals, and their definitive transformation into
what is called the _monopetalous_ corolla. The term _monopetaloid_
ought then to be rejected, if we are to believe that the
_monopetalous_ corolla is the result of the metamorphosis of a
single leaf. The word _gamopetalous_, or, rather, _gamophyllous_,
is preferable. As we have remarked in reference to the calyx, we
shall here repeat that the expressions _bilobed_, _trilobed_,
_quadrilobed_, or _bipartite_, _tripartite_, _quadripartite_
corollas, are radically vicious, because they are the consequence of
a false point of view, according to which the _monopetalous_ corolla
will be simply a single metamorphosed leaf, susceptible of being more
or less deeply divided from top to bottom.
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