Some of the admissions made in this paragraph have an important bearing
on the theory of natural selection. Far from supposing that fortuitous
destruction would have no influence on the course of natural selection,
it can be shown that it would have a most disastrous effect. In many
cases the destruction comes in the form of a catastrophe to the
individuals, so that small differences in structure, whether
advantageous or not, are utterly unavailing. Our experience shows us
that a destruction of this sort is going on around us all the time, and
accounts in large part for the way in which the majority of animals and
plants are destroyed. Unless, for example, a seed happen to fall on a
place suitable for its growth, it will perish without respect to a
slight advantage it may have over other seeds of its kind. Of the
thousands of eggs laid by one starfish, chance alone will decide whether
one or another embryo is destroyed by larger animals, or if they escape
this danger, the majority of them may be carried out to sea, where it
will not be of the least avail if one individual has a slight advantage
over the others. Darwin admits this, but adds that, if only a thousandth
part is developed, yet of those that do survive the best adapted
individuals will tend to propagate their kind in larger numbers than the
less well adapted. The argument is not, however, so simple as it appears
to be on the surface. I pass over, for the present, the apparent
inconsequence in this statement that the best adapted individuals will
tend to propagate their kind in larger numbers. It is not by any means
certain that this is the case. Darwin’s meaning is, however, fairly
clear, and can be interpreted to mean this: after the fortuitous
destruction has finished, there will be a further competition of the
survivors amongst themselves and with the surrounding conditions. In
this higher competition, which is less severe, small individual
differences suffice to determine the survival of certain individuals.
These are, therefore, selected.
In this argument it is assumed that a second competition takes place
after the first destruction of individuals has occurred, and this
presupposes that more individuals reach maturity than there is room for
in the economy of nature. But we do not know to what extent this takes
place. If only as many mature as can survive, then the second
competition does not take place. If, on the other hand, fewer mature
than there is room for, then again competition does not take place. And
if at all times selection is not rigorously carried out, everything may
be lost that has been so laboriously gained. We see then that the result
that Darwin imagines would take place, can be carried out only when more
individuals reach maturity than there is room for (if it is a case of
competition with one another), or that escape their enemies (if it is a
question of competition with other forms).
Public-domain text, read in full here on John Shaqi.
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