Weismann finds the solution in the method by which the breeder has
obtained his results in artificial selection. For instance, the
long-tailed variety of the domestic cock of Japan owes its existence, it
is claimed, to skilful selection, and not at all to the circumstance
that, at some period of the race’s history, a cock with tail-feathers
six feet in length suddenly and spasmodically appeared.
Weismann continues: “Now what does this mean? Simply that the hereditary
diathesis, the germinal constitution (the _Anlage_) of the breed was
changed in the respect in question, and our conclusion from this and
numerous similar facts of artificial selection runs as follows: _by the
selection alone of the plus or minus variations of a character is the
constant modification of that character in the plus or minus direction
determined_. Obviously the hereditary _diminution_ of a part is also
effected by the simple selection of the individuals in each generation
possessing the smallest parts, as is proved, for example, by the tiny
bills and feet of numerous breeds of doves. We may assert, therefore, in
general terms: a definitely directed progressive variation of a given
part is produced by continued selection in that definite direction. This
is no hypothesis, but a direct inference from the facts and may also be
expressed as follows: _by a selection of the kind referred to the germ
is progressively modified in a manner corresponding with the production
of a definitely directed progressive variation of the part_.”
So far there is nothing essentially new offered, since Darwin often
tacitly recognized that the standard of variation could be raised in
this way, and in some places he has made definite statements that this
will take place. Weismann thinks that after each selection, fluctuation
will then occur around a higher average (mode). He says “that this is a
fact,” and is proved by the case of the Japanese cock. It need scarcely
be pointed out that it is an assumption, based on what is supposed to
have taken place in this bird, and is not a “fact.”
Public-domain text, read in full here on John Shaqi.
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