Spencer claims that, since 1886, when he first published this argument,
nothing like an adequate response has been made; and I think he might
have added that an adequate answer is not likely to be forthcoming,
since nothing short of a demonstration of how the giraffe really evolved
is likely to be considered as sufficient. Wallace’s reply, that the
changes in question could have been brought about by natural selection,
since similar changes have been brought about by artificial selection,
is regarded as inadequate by Spencer, since it assumes a parallel which
does not exist. Nevertheless, Wallace’s reply contains, in my opinion,
the kernel of the explanation, in so far as it assumes that congenital
variation[19] may suffice to account for the origin of a form even as
bizarre as that of the giraffe. The ancon ram and the turnspit dog were
marked departures from the normal types, and yet their parts were
sufficiently coördinated for them to carry out the usual modes of
progression. It would not have been difficult, if we adopted Spencer’s
mode of arguing, to show that these new forms could not possibly have
arisen as the result of congenital variations.
Footnote 19:
Wallace assumes fluctuating variation to suffice, but in this I cannot
agree with him.
Again, it might be argued that the large, powerful dray-horse could not
have arisen through a series of variations from the ordinary horse,
because, even if variations in the right direction occurred in the
fore-quarters, it is unlikely that similar variations would occur in the
hind-quarters, etc. Yet the feat has been accomplished, and while it is
difficult to prove that the inheritance of acquired characters has not
had a hand in the process, it is improbable that this has been the case,
but rather that artificial selection of some kind of variations has been
the factor at work.
So long as the Lamarckian theory is supported by arguments like these,
it can never hope to be established with anything more than a certain
degree of probability. If it is correct, then its demonstration must
come from experiment. This brings us to a consideration of the
experimental evidence which has been supposed by some writers to give
conclusive proof of the validity of the theory.
Public-domain text, read in full here on John Shaqi.
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