join the tops of these perpendiculars, the curve of frequency is the
result.
[Illustration:
Fig. 3.—Curves of frequency, etc.
A, normal curve.
B, showing the method of arranging individuals in lines containing
similar kinds of individuals.
C, curve that is skew to the right.
D, polygon of frequencies of horns of rhinoceros beetles.
(After Davenport.)]
“In arranging the individuals it will be found, as has been said, that
certain groups contain more individuals. They will form the longest
line. This value that occurs with the greatest frequency is called the
mode. The position of this modal class in the polygon is one of the
points of importance, and the spread of the polygon at its base is
another. A polygon with a low mode and a broad range means great
variability. The range may, however, be much affected by a single
individual standing far removed from the rest, so that a polygon
containing such an individual might appear to show greater variation
than really exists. Therefore we need a measure of variability that
shall take into account the departures of all the individuals from the
mode. One such measure is the arithmetical average of all the departures
from the mean in both directions; and this measure has been widely
employed. At present another method is preferred, namely, the square
root of the squared departures. This measure is called the standard
deviation. The standard deviation is of great importance, because it is
the index of variability.”[21]
Footnote 21:
Davenport, C. B., “The Statistical Study of Biological Problems,”
_Popular Science Monthly_, September, 1900.
Of the different kinds of polygons there are two main sorts, the simple
and the complex. The former have only a single mode, the latter have
more than one mode. Some simple polygons lie symmetrically on each side
of the mode, Figure 3 A; others are unsymmetrical or skew, Figure 3 B.
The skew polygon generally extends out on one side farther than on the
other. It has been suggested that when a polygon is symmetrical the
species is not changing, and when skew that the species is evolving in
the direction of the longer base. This assumes that the sort of
variation measured by these curves is of the kind of which evolution is
made up, but this is a question that we must further consider. How far
the change indicated by the skew curve may be carried is also another
point for further examination.
A complex polygon of variation, Figure 3 D, has been sometimes
interpreted to mean that two subgroups exist in a species, as is well
shown in the case of the rhinoceros beetle described by Bateson. Two
kinds of male individuals exist, some with long horns, others with short
horns; each with a mode of its own, the two polygons overlapping. Other
complex polygons may be due to changes occurring at different times in
the life of the individual, as old age, for example.
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