Pearson believes that “if a race has been long under the same
environment it is probable that only periodic selection is at work,
maintaining its stability. Change the environment and a secular change
takes place, the deviations from the mode previously destroyed giving
the requisite material.” “Clearly periods of rapidly changing
environment, of great climatological and geological change, are likely
to be associated with most marked secular selection. To show that there
is little or no change year by year in the types of rabbit and wild
poppy in our English fields, or of daphnia in our English ponds, is to
put forward no great argument for the inefficiency of natural selection.
Take the rabbit to Australia, the wild poppy to the Cape, the daphnia
into the laboratory, and change their temperature, their food supply,
and the chemical constituents of water and air, and then the existence
of no secular selection would indeed be a valid argument against the
Darwinian theory of evolution.” In regard to the last point, it should
be noted that, even if under the changed conditions a change in the mode
took place, as Pearson assumes, it does not follow necessarily that
selection has had anything to do with it, but the environment may have
directly changed the forms. Furthermore, and this is the essential
point, even if selection does act to the extent of changing the mode, we
should not be justified in concluding that this sort of change could go
on increasing as long as the selection lasts. All that might happen
would be to keep the species up to the highest point to which
fluctuating variation can be held. This need not lead to the formation
of new species, or direct the course of evolution.
Pearson points out further that, even if we suppose that a secular
change is produced in a new environment, we cannot explain how species
may break up into two or more races that are relatively infertile.
Suppose two groups of individuals, subjected to different environments,
become isolated geographically. Two local races will be produced.
“Isolation may account for the origin of local races, but never for the
origin of species unless it is accompanied by a differential fertility.”
In other words, Pearson thinks that, unless the reproductive organs are
correlated with other organs, in such a way that as these organs change
the interracial fertility of the germ-cells is altered, so that in the
two changed groups the individuals are no longer interfertile, new
species cannot be accounted for, since their mutual infertility is one
of their most characteristic features. “Without a barrier to
intercrossing during differentiation the origin of species seems
inexplicable.”
Public-domain text, read in full here on John Shaqi.
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