Footnote 24:
Bateson, in his book on “Mendel’s Principles of Heredity,” has given
an admirable presentation of Mendel’s results. I have relied largely
on this in my account.
The importance of Mendel’s results and their wide application is
apparent from the results in recent years of De Vries, Correns,
Tschermak, Bateson, Castle, and others. Mendel carried out his
experiments on the pea, _Pisum sativum_. Twenty-two varieties were used,
which had been proven by experiment to be pure breeds. When crossed they
gave perfectly fertile offspring. Whether they all have the value of
varieties of a single species, or are different subspecies, or even
independent species, is of little consequence so far as Mendel’s
experiments are concerned. The flower of the pea is especially suitable
for experiments of this kind. It cannot be accidentally fertilized by
foreign pollen, because the reproductive organs are inclosed in the keel
of the flower, and, as a rule, the anthers burst and cover the stigma of
the same flower with its own pollen before the flower opens. In order to
cross-fertilize the plants it is necessary to open the young buds before
the anthers are mature and carefully remove all the anthers. Foreign
pollen may be then, or later, introduced.
The principle involved in Mendel’s law may be first stated in a
theoretical case, from which a certain complication that appears in the
actual results may be removed.
If _A_ represent a variety having a certain character, and _B_ another
variety in which the same character is different, let us say in color,
and if these two individuals, one of each kind, are crossed, the hybrid
may be represented by _H_. If a number of these hybrids are bred
together, their descendants will be of three kinds; some will be like
the grandparent, _A_, in regard to the special character that we are
following, some will be like the other grandparent, _B_, and others will
be like the hybrid parent, _H_. Moreover, there will be twice as many
with the character _H_, as with _A_, or with _B_.
A B
↘ ↙
H
↙|↘
A | B
↙ | ↘
A | B
H
↙|↘
A | B
↙ | ↘
A | B
H
↙|↘
A | B
H
If now we proceed to let these _A_’s breed together, it will be found
that their descendants are all _A_, forever. If the _B_’s are bred
together they produce only _B_’s. But when the _H_’s are bred together
they give rise to _H_’s, _A_’s, and _B_’s, as shown in the accompanying
diagram. In each generation, the _A_’s will also breed true, the _B_’s
true, but the _H_’s will give rise to the three kinds again, and always
in the same proportion.
Thus it is seen that the hybrid individuals continue to give off the
pure original forms, in regard to the special character under
consideration. The numerical relation between the numbers is also a
striking fact. Its explanation is, however, quite simple, and will be
given later.
Public-domain text, read in full here on John Shaqi.
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