From the evidence that his results have given, De Vries believes it is
probable that mutation has occurred in all directions. In the same way
that Darwin supposed that individual or fluctuating variations are
scattering, so also De Vries believes that the new forms that arise
through mutation are scattering. On this point it seems to me that De
Vries may be too much prejudiced by his results with the evening
primrose. If, as he supposes, many forms, generally ranked as varieties,
are really elementary species, it seems more probable that the mutation
of a form may often be limited to the production of one or of only a
very few new forms. The single variations, or sports, point even more
strongly in favor of this interpretation. Moreover, the general problem
of evolution from a purely theoretical point of view is very much
simplified, if we assume that the kinds of mutating forms may often be
very limited, and that mutations may often continue to occur in a direct
line. On this last point, De Vries argues that the evidence from
paleontology cannot be trusted, for all that we can conclude from fossil
remains is that certain mutations have dominated, and have been
sufficiently abundant to leave a record. In other words, the conditions
may have been such that only certain forms could find a foothold.
De Vries asks whether there are for each species periods of mutation
when many and great changes take place, and periods when relatively
little change occurs. The evidence upon which to form an opinion is
scanty, but De Vries is inclined to think that such periods do occur. It
is at least certain from our experience that there are long periods when
we do not see new forms arising, while at other times, although we know
very few of them, epidemics of change may take place. The mutative
period which De Vries found in the evening primrose is the best-known
example of such a period of active mutation. Equally important for the
descent theory is the idea that the same mutation may appear time after
time. There is good evidence to show that this really occurs, and in
consequence the chances for the perpetuation of such a form are greatly
increased. Delbœuf, who advocated this idea of the repeated reappearance
of a new form, has also attempted to show that if this occurs the new
form may become established without selection of any kind taking
place,—the time required depending upon the frequency with which the new
form appears. This law of Delbœuf, De Vries believes, is correct from
the point of view of the mutation theory. It explains, in a very simple
way, the existence of numerous species-characters that are entirely
useless, such, for instance, as exist between the different elementary
species of _Draba verna_. “According to the selection theory only useful
characters can survive; according to the mutation theory, useless
characters also may survive, and even those that may be hurtful to a
small degree.”
Public-domain text, read in full here on John Shaqi.
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