In regard to the song-sparrow (_Melospiza_), Coues writes: “The type of
the genus is the familiar and beloved song-sparrow, a bird of constant
characters in the east, but in the west is split into numerous
geographical races, some of them looking so different from typical
_fasciata_ that they have been considered as distinct species, and even
placed in other genera. This differentiation affects not only their
color, but the size, relative proportions of parts, and particularly the
shape of the bill; and it is sometimes so great, as in the case of _M.
cinerea_, that less dissimilar looking birds are commonly assigned to
different genera. Nevertheless the gradation is complete, and affected
by imperceptible degrees.... The several degrees of likeness and
unlikeness may be thrown into true relief better by some such
expressions as the following, than by formal antithetical phrases: (1)
The common eastern bird commonly modified in the interior into the
duller colored (2) _fallax_. This in the Pacific watershed, more
decidedly modified by deeper coloration,—broader black streaks in (3)
_hermanni_, with its diminutive local race (4) _samuelis_, and more
ruddy shades in (5) _guttata_ northward, increasing in intensity with
increased size in (6) _rafina_. Then the remarkable (7) _cinerea_,
insulated much further apart than any of the others. A former American
school would probably have made four ‘good species,’ (1) _fasciata_, (2)
_samuelis_, (3) _rafina_, (4) _cinerea_.”
Somewhat similar relations are found in three other genera of finches.
Thus Passerella is “imperfectly differentiated”; Junco is represented by
one eastern species, but in the west the stock splits up into numerous
forms, “all of which intergrade with each other and with the eastern
bird. Almost all late writers have taken a hand at Junco, shuffling them
about in the vain attempt to decide which are ‘species’ and which
‘varieties.’ All are either or both, as we may elect to consider them.”
In the distribution of the genus Pipilo similar relations are found.
There is an eastern form much more distinct from the western forms than
these are from each other.
Public-domain text, read in full here on John Shaqi.
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