Let us consider now the more general questions involved in these
symmetrical and asymmetrical relations between the organism and its
environment. In what sense, it may be asked, is the symmetry of a form
an adaptation to its environment? That the kind of symmetry gives to the
animal in many cases a certain advantage in relation to its environment
is so evident that I think it will not be questioned. The main question
is how this relation is supposed to have been attained. Three points of
view suggest themselves: First, that the form has resulted directly from
the action of the environment upon the organism. This is the Lamarckian
point of view, which we rejected as improbable. Second, that the form
has been slowly acquired by selecting those individual variations that
best suited it to a given set of surrounding conditions. This is the
Darwinian view, which we also reject. The third, that the origin of the
form has had nothing to do with the environment, but appeared
independently of it. Having, however, appeared, it has been able to
perpetuate itself under certain conditions.
It should be pointed out that the Darwinian view does not suppose that
the environment actually produces any of the new variations which it
selects after they have appeared, but in so far as the environment
selects individual differences it is supposed to determine the direction
in which evolution takes place. On the theory that evolution has taken
place independently of selection, this latter is not supposed to be the
case; the finished products, so to speak, are offered to the
environment; and if they pass muster, even ever so badly, they may
continue to propagate themselves.
The asymmetrical form of certain animals living in a symmetrical
environment might be used as an argument to show that the relation of
symmetry between an animal and its environment can easily be overstepped
without danger. The enormous claw of the fiddler-crab must throw the
animal out of all symmetrical relation with its environment, and yet the
species flourishes. The snail carries around a spiral hump that is
entirely out of symmetrical relation with the surroundings of a snail.
These facts, few though they are, yet suffice to show, I believe, that
the relation of symmetry between the organism and its environment may
be, and is no doubt in many cases, more perfect than the requirements of
the situation demand. The fact that animals made unsymmetrical through
injuries (as when a crab loses several legs on one side, or a worm its
head) can still remain in existence in their natural environment, is in
favor of the view that I have just stated. By this I do not mean to
maintain that a symmetrical form does not have, on the whole, an
advantage over the same form rendered asymmetrical, but that this
relation need not have in all forms a selective value, and if not, then
it cannot be the outcome of a process of natural selection.
Public-domain text, read in full here on John Shaqi.
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