Many of the facts as to the occurrence of these cases were known
before Darwin worked on them; but very little had been ascertained in
regard to the sexual relation between the dimorphic and trimorphic
forms, and it was here that Darwin obtained his most interesting
results.
The outcome of these different crossings are most curious. In the table,
page 364, the results of the four combinations are given. It will be
seen at once that the legitimate unions give more capsules, and the
seeds weigh more, than in the illegitimate unions.
The behavior of the offspring from seeds of legitimate and
illegitimate origin is even more astonishing. Darwin found in _Primula
veris_ (the form just described) that the seeds from the short-styled
form fertilized with pollen from the same form germinated so badly
that he obtained only 14 plants, of which 9 were short-styled and 5
long-styled. The long-styled form fertilized with its own-styled
pollen produced “in the first generation 3 long-styled plants. From
their seed 53 long-styled grandchildren were produced; from their seed
4 long-styled great-grandchildren; from their seed 20 long-styled
great-great-grandchildren; and lastly, from their seed 8 long-styled
and 2 short-styled great-great-great-grandchildren.”
══════════════╤═════════╤═════════╤═════════╤═════════╤═════════
│Number of│ │ Maximum │ Minimum │ Average
Nature of │ Flowers │Number of│of Seeds │of Seeds │ No. of
Union │Fertilized│ Seed │ in any │ in any │Seeds per
│ │Capsules │ one │ one │ Capsule
│ │ │ Capsule │ Capsule │
──────────────┼─────────┼─────────┼─────────┼─────────┼─────────
Long-styled │ 10 │ 6 │ 62 │ 34 │ 46.5
form by │ │ │ │ │
pollen of │ │ │ │ │
short-styled│ │ │ │ │
form: │ │ │ │ │
Legitimate │ │ │ │ │
union. │ │ │ │ │
──────────────┼─────────┼─────────┼─────────┼─────────┼─────────
Long-styled │ 20 │ 4 │ 49 │ 2 │ 27.7
form by │ │ │ │ │
own-form │ │ │ │ │
pollen: │ │ │ │ │
Illegitimate│ │ │ │ │
union. │ │ │ │ │
──────────────┼─────────┼─────────┼─────────┼─────────┼─────────
Short-styled │ 10 │ 8 │ 61 │ 37 │ 47.7
form by │ │ │ │ │
pollen of │ │ │ │ │
long-styled │ │ │ │ │
form: │ │ │ │ │
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