Hertwig has recently raised some new points of issue in regard to the
recapitulation theory, and since he may appear to have penetrated
farther than most other embryologists of the present time, it will be
necessary to examine his view somewhat carefully. He speaks of the
germ-cell (egg, or spermatozoön) as a species-cell, because it contains,
in its finer organization, the essential features of the species to
which it belongs. There are as many of these kinds of cells as there are
different kinds of animals and plants. Since the bodies of the higher
animals have developed from these species-cells, so the latter must have
passed in their phylogeny through a corresponding development from a
simple to a more and more complex cell-structure. “Our doctrine is, that
the species-cell, even as the adult, many-celled representative of the
species, has passed through a progressive, and, indeed, in general a
corresponding development in the course of phylogeny. This view appears
to stand in contradiction to the biogenetic law. According to the
formula that Haeckel has maintained, the germ development is an epitome
of the genealogy; or the ontogeny is a recapitulation of the phylogeny;
or, more fully, the series of forms through which the individual
organism passes during its development from the egg-cell to the finished
condition is a short, compressed repetition of the longer series of
forms which the forefathers of the same organism, or the stem-form of
the species, has passed through, from the earliest appearance of
organisms to the present time.” “Haeckel admits that the parallel may be
obliterated, since much may be absent in the ontogeny that formerly
existed in the phylogeny. If the ontogeny were complete, we could trace
the whole ancestry.” Hertwig states further, that “The theory of
biogenesis[12] makes it necessary to change Haeckel’s expression of the
biogenetic law, so that a contradiction contained in it may be removed.
We must drop the expression ‘repetition of the form of extinct
forefathers,’ and put in its place the repetition of forms which are
necessary for organic development, and lead from the simple to the
complex. This conception may be illustrated by the egg-cell.”
Footnote 12:
This term, by which Hertwig designates a particular view of his own,
has been already preoccupied in a much wider sense by Huxley to mean
that all life comes from preëxisting life. Hertwig means by the theory
of biogenesis that as the egg develops there is a constant interchange
between itself and its surroundings.
Public-domain text, read in full here on John Shaqi.
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