Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
1895. _Pappogeomys_ Merriam, N. Amer. Fauna, 8:145, January 31.
1895. _Cratogeomys_ Merriam, N. Amer. Fauna, 8:150, January 31.
Type: _Geomys merriami_ Thomas.
1895. _Platygeomys_ Merriam, N. Amer. Fauna, 8:162, January 31.
Type: _Geomys gymnurus_ Merriam; Hooper, Jour. Mamm.,
27:397, November 25, 1946.
_Type._--_Geomys bulleri_ Thomas, 1892, from near Talpa, west slope
Sierra de Mascota, 8500 ft. (actually about 5000 ft.), Jalisco.
_Chronologic range._--Late Pliocene, from deposits of early Blancan
age (Benson local fauna, Arizona) to the Recent. However in the
Pleistocene, only late Pleistocene records are known, and
_Pappogeomys_ has not been found in early (late Blancan) or middle
(Irvingtonian) Pleistocene local faunas. Presumably the genus was
restricted to México during the Pleistocene until post-Wisconsin time.
_Description and discussion._--The size ranges from as little as in
the smaller kinds of _Thomomys_ to the maximum attained in the
subfamily and matched elsewhere perhaps in only a few of the larger
subspecies of _Orthogeomys grandis_. Depending on the species and
subgenus, the form of the skull varies from generalized to
specialized. The generalized skulls are short and not especially
narrow; the zygomatic arches are spread laterally so far that the
breadth across them exceeds the breadth across the mastoid processes.
The most specialized skulls are platycephalic and the breadth across
the mastoid processes equals or exceeds the breadth across the
zygomatic arches (even so, the zygomatic arches are still relatively
widespread). In correlation with the great breadth of the posterior
part of the cranium, the rami of the mandibles diverge widely
posteriolaterally and the angular processes are remarkably elongated.
The rostrum is moderately broad in most species, but not nearly so
broad and heavy as in _Orthogeomys_.
The single deep, median sulcus on the outer surface of the upper
incisor is slightly displaced to the inner side of the tooth. The
posterior surface of P4 lacks enamel (small vestige found on lingual
end of posterior wall in only two adult individuals--UA 3260 and KU
100442, of the subgenus _Pappogeomys_); the other three plates are
fully developed as usual. The p4 is provided with four fully developed
enamel plates, in the pattern characteristic of the tribe Geomyini. In
the p4 of the late Pliocene species (_P. bensoni_) the re-entrant
angles are open (obtuse), a trait that is evidently primitive in the
Geomyini.
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