Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
The occlusal pattern, at least in M1 and M2, in the final stages of
wear in _Dikkomys_ resembles that in the subfamily Entoptychinae, but
the U-pattern develops on only the first and probably the second molar
in _Dikkomys_ and not on all of the cheek teeth as it does in the
entoptychines. Judging from the material that has been described, the
U-pattern did not develop in the lower teeth of _Dikkomys_ until the
Hemingfordian (_D. woodi_), upper Rosebud, and specimens of _D.
matthewi_ from the earlier Arikareean, lower Harrison, suggest that
the modified H-pattern, with secondary coalescence at the edge of the
protomeres, persisted throughout life, without developing the
U-pattern in the final stages of wear.
Essentially the same patterns of wear characterize the genus
_Pliosaccomys_, except that the earlier stages were telescoped and the
second stage was omitted while another (final) stage was added. The
stages are reconstructed in sequence in figure 4, and all are based on
preserved dentitions, as follows:
(1) The first phases of wear produced the pattern (Fig. 4E and I)
described for _Dikkomys_ in the previous account (Fig 4A).
(2) A small additional amount of wear produced the 2nd stage (Fig. 4F
and J) characterized by a U-pattern, formed by union of the anterior
and posterior columns at the edge of the protomeres of the first and
second molars, both above and below, without first forming an H-shaped
pattern. Union at the mid-points thus was omitted from the sequence of
wear in these two teeth. In the premolars and third molars the
primitive H-pattern did form, as in _Dikkomys_. The pattern of wear in
the first two molars is the same as in the entoptychines of the early
Miocene. The trend of evolution through which the _Pliosaccomys_
lineage passed must have featured a progressively earlier union at the
edge of the tooth until the lateral coalescence occurred
simultaneously with the median union. At that stage, emphasis was
shifted to the union at the edge of the tooth, and eventually the
teeth failed to unite at their mid-points and the U-pattern developed
directly. Therefore, the horizontally deep re-entrant fold that
separates the two lophs of the U-pattern is equivalent to one fold
plus the apex of the opposite fold.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account