_Geomys_, slightly more specialized than _Zygogeomys_, must also be regarded as one of the most primitive of the living genera. Primitive features that have been retained are the generalized type of skull, the bisulcate pattern of grooves on the upper incisor, and the retention of enamel plates on both the anterior and posterior walls of M1 and M2 (see Fig. 9A). All of these primitive features are shared with _Zygogeomys_. In addition, three other trends, or specializations, in evolution characterize the phyletic development of _Geomys_. One major trend is toward loss of the enamel plate from the posterior wall of P4. No trace of enamel remains on the posterior wall of this tooth in late Pleistocene or Recent species of _Geomys_, and at least one of the earlier species (_quinni_) was also characterized by loss of this enamel plate. Secondly, M3 retains only a vestige of the primitive bicolumnar pattern after the initial stages of wear. In most Recent specimens, especially of the species _G. bursarius_, the lateral re-entrant fold and the heel of M3 are small, and the re-entrant inflection is hardly evident. The lateral fold is more frequently well-developed in Irvingtonian species than in living species (White and Downs, 1961:13), illustrating progressive loss of the bicolumnar pattern in Pleistocene evolution. A third trend involves the modification of the lateral folds of the premolars. Primitively the angles of these folds are broadly open or divergently V-shaped, and some of the earliest species of _Geomys_, for example _G. quinni_, have retained this feature throughout life. Nevertheless, the main trend is toward progressive compression of the folds resulting in their walls being more nearly perpendicular, and parallel, to the long axis of the tooth. Obtuse re-entrant angles persist in premolars of young individuals of Irvingtonian species, but the adults are characterized by well-compressed folds, as in Recent species.