At the present time we are incomparably more favourably placed than
Darwin was for answering this question of all questions. We have at
our command an incomparably greater wealth of material than he had at
his disposal. And we are more fortunate than he in this respect, that
we now know transition-forms which help to fill up the gap, still
great, between the lowest human races and the highest apes. Let us
consider for a little the more essential additions to our knowledge
since the publication of _The Descent of Man_.
Since that time our knowledge of animal embryos has increased
enormously. While Darwin was obliged to content himself with comparing
a human embryo with that of a dog, there are now available the
youngest embryos of monkeys of all possible groups (Orang, Gibbon,
Semnopithecus, Macacus), thanks to Selenka's most successful tour in
the East Indies in search of such material. We can now compare
corresponding stages of the lower monkeys and of the Anthropoid apes
with human embryos, and convince ourselves of their great resemblance
to one another, thus strengthening enormously the armour prepared by
Darwin in defence of his view on man's nearest relatives. It may be
said that Selenka's material fills up the blanks in Darwin's array of
proofs in the most satisfactory manner.
The deepening of our knowledge of comparative anatomy also gives us much
surer foundations than those on which Darwin was obliged to build. Just of
late there have been many workers in the domain of the anatomy of apes and
lemurs, and their investigations extend to the most different organs. Our
knowledge of fossil apes and lemurs has also become much wider and more
exact since Darwin's time: the fossil lemurs have been especially worked up
by Cope, Forsyth Major, Ameghino, and others. Darwin knew very little about
fossil monkeys. He mentions two or three anthropoid apes as occurring in
the Miocene of Europe,[111] but only names _Dryopithecus_, the largest form
from the Miocene of France. It was erroneously supposed that this form was
related to _Hylobates_. We now know not only a form that actually stands
near to the gibbon (_Pliopithecus_), and remains of other anthropoids
(_Pliohylobates_ and the fossil chimpanzee, _Palaeopithecus_), but also
several lower catarrhine monkeys, of which _Mesopithecus_, a form nearly
related to the modern Sacred Monkeys (a species of _Semnopithecus_) and
found in strata of the Miocene period in Greece, is the most important.
Quite recently, too, Ameghino's investigations have made us acquainted with
fossil monkeys from South America (_Anthropops_, _Homunculus_), which,
according to their discoverer, are to be regarded as in the line of human
descent.
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