The operation of these principles cannot be denied. Man is still an animal, subject to zoological as well as mechanical laws. The dark influence of heredity continues to be effective; and psychical development had begun in lower organic forms,--perhaps with life itself. The organic and the social struggles for existence are manifestations of the same principle. Environment and climatic influence must be called in to explain not only the differentiation of the great racial sections of humanity, but also the varieties within these sub-species and, it may be, the assimilation of distinct varieties. Ritter's _Anthropogeography_ has opened a useful line of research. But on the other hand, it is urged that, in explaining the course of history, these principles do not take us very far, and that it is chiefly for the primitive ultra-prehistoric period that they can account for human development. It may be said that, so far as concerns the actions and movements of men which are the subject of recorded history, physical environment has ceased to act mechanically, and in order to affect their actions must affect their wills first; and that this psychical character of the causal relations substantially alters the problem. The development of human societies, it may be argued, derives a completely new character from the dominance of the conscious psychical element, creating as it does new conditions (inventions, social institutions, etc.) which limit and counteract the operation of natural selection, and control and modify the influence of physical environment. Most thinkers agree now that the chief clews to the growth of civilisation must be sought in the psychological sphere. Imitation, for instance, is a principle which is probably more significant for the explanation of human development than natural selection. Darwin himself was conscious that his principles had only a very restricted application in this sphere, as is evident from his cautious and tentative remarks in the 5th chapter of his _Descent of Man_. He applied natural selection to the growth of the intellectual faculties and of the fundamental social instincts, and also to the differentiation of the great races or "sub-species" (Caucasian, African, etc.) which differ in anthropological character.[241]