It will be sufficient to indicate an essential difference in the
process of development in the higher and lower Crustacea. In the latter
all new body-segments and limbs which insert themselves between the two
terminal regions of the Nauplius, are formed in uninterrupted sequence
from before backwards; in the former there is further a new formation
in the middle of the body (the middle-body), which pushes itself in
between the fore-body and the abdomen in the same way, as these have
done on their part between the head and tail of the Nauplius. Thus,
that which appears probable even from the comparison of the limbs of
the adult animal, finds fresh support in the developmental history,
namely, that the lower Crustacea, like the Insects, are entirely
destitute of the region of the body corresponding to the middle-body of
the Malacostraca. It seems probable that the swimming feet of the
Copepoda, as also of the pupæ of Cirripedia and Rhizocephala, represent
the abdominal feet of the Malacostraca, that is to say, are derived by
inheritance from the same source with them.
It would be easy to weave together the separate threads furnished by
the young forms of the various Crustacea, into a general picture of the
primitive history of this class. Such a picture, drawn with a little
skill, and finished in lively colours, would certainly be more
attractive than the dry discussions which I have tacked on to the
developmental history of these animals. But the mode of weaving in the
loose threads would still in many cases be arbitrary, and to be
effected with equal justice in various ways; and many gaps would still
have to be filled up by means of more or less bold assumptions. Those
who have not wandered much in this region of research would then
readily believe that they were standing upon firm ground, where mere
fancy had thrown an airy bridge; those acquainted with the subject, on
the other hand, would soon find out these weak points in the structure,
but would then be easily led to regard even what was founded upon well
considered facts, as merely floating in the air. To obviate these
misconceptions of its true contents from either side, it would be
necessary to accompany such a picture throughout with lengthy, dry
explanations. This has deterred me from further filling in the outline
which I had already sketched.
I will only give, as an example, the probable history of the production
of a single group of Crustacea, and indeed of the most abnormal of all,
the RHIZOCEPHALA, which in the sexually mature state differ so
enormously even from their nearest allies, the Cirripedia, and from
their peculiar mode of nourishment stand quite alone in the entire
animal kingdom.
Public-domain text, read in full here on John Shaqi.
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