Plants -- Identification; Wild flowers -- Great Britain
The peculiar arrangement of the stamens above described is sufficient in
itself to distinguish the genus _Salvia_ from all the other Labiates,
and the importance of the peculiarity in connexion with the pollination
of the flower is so interesting that we may well spend a few minutes in
studying it before passing on to other species. In the first place it
should be mentioned that the stamens of _Salvia_ are mature before the
stigma, and that, as a consequence, self-pollination is impossible. The
lower, abortive anthers of the two stamens are joined together and form
a little valve which closes the throat of the corolla tube. Each one,
however, has a notch in its inner side, and the two notches, meeting in
the middle, form a little hole. When a bee visits the flower, it alights
on the lower lip of the corolla, and thrusts its tongue through the
hole to reach the nectar at the base of the tube. In doing this it
pushes the abortive anthers backwards, and the upper, fertile anther
cells, which rest under the arched upper lip, are thus made to swing
downwards and forwards so that they touch the bee's back; and, if they
are ripe, to deposit some pollen. After the pollen has been thus
removed, the style lengthens and curves downward, bringing the stigma,
which is now mature, to the position previously occupied by the fertile
anther cells at the time they were made to swing downwards by the bee.
Thus, if a bee which has previously visited a flower with mature anthers
now comes to one in which the stigma is ripe, the pollen dusted on its
back by the former is rubbed against the stigma of the latter, and
cross-fertilisation is the result.
[Illustration: SECTION OF THE FLOWER OF _Salvia_.
1. Stigma--not yet mature. 2. Stamen.]
[Illustration: THE SELF-HEAL.]
The Common Marjoram (_Origanum vulgare_) is an aromatic plant that often
grows in great abundance on dry hilly pastures, especially in limestone
and chalky districts. Its stem is thin and hairy, a foot or more in
height; and the leaves are stalked, ovate, blunt, slightly toothed,
downy, and about an inch in length. The flowers, which bloom from July
to September, are of a rosy purple colour, in numerous globular
clusters, the whole inflorescence forming a leafy panicle. The
overlapping bracts are about as long as the calyx, and usually tinged
with red or purple; the calyx has five, short, equal teeth, and is very
hairy in the throat; the corolla is about twice the length of the
calyx, and has four lobes, the upper of which is a little broader than
the others; and the stamens are in two pairs, one pair longer than the
other. It will be noticed that some of the flowers are larger than
others, and that these are perfect, while the smaller ones have no
stamens.
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