The simplest kind of fruit is a _ripened 1-loculed ovary_. The first
stage in complexity is a ripened _2- or many-loculed ovary_. Very
complex forms may arise by the _attachment of other parts to the
ovary_. Sometimes the style persists and becomes a beak (mustard pods,
dentaria, Fig. 224), or a tail as in clematis; or the calyx may be
attached to the ovary; or the ovary may be embedded in the receptacle,
and ovary and receptacle together constitute the fruit: or an involucre
may become a part of the fruit, as possibly in the walnut and hickory
(Fig. 225), and cup of the acorn (Fig. 226). The chestnut and the beech
bear a prickly involucre, but the nuts, or true fruits, are not grown
fast to it, and the involucre can scarcely be called a part of the
fruit. A ripened ovary is a =pericarp=. A pericarp to which other parts
adhere has been called an accessory or reënforced fruit. (Page 169.)
[Illustration: FIG. 225.--HICKORY-NUT. The nut is the fruit, contained
in a husk.]
[Illustration: FIG. 226.--LIVE-OAK ACORN. The fruit is the “seed” part;
the involucre is the “cup.”]
[Illustration: FIG. 227.--KEY OF SUGAR MAPLE.]
[Illustration: FIG. 228.--KEY OF COMMON AMERICAN ELM.]
Some fruits are =dehiscent=, or split open at maturity and liberate the
seeds; others are =indehiscent=, or do not open. A dehiscent pericarp
is called a =pod=. The parts into which such a pod breaks or splits
are known as =valves=. In indehiscent fruits the seed is liberated by
the decay of the envelope, or by the rupturing of the envelope by the
germinating seed. Indehiscent winged pericarps are known as =samaras=
or =key fruits=. Maple (Fig. 227), elm (Fig. 228), and ash (Fig. 93)
are examples.
=Pericarps.=--The simplest pericarp is a dry, one-seeded, indehiscent
body. It is known as an =akene=. A head of akenes is shown in Fig.
229, and the structure is explained in Fig. 230. Akenes may be seen in
buttercup, hepatica, anemone, smartweed, buckwheat.
[Illustration: FIG. 229.--AKENES OF BUTTERCUP.]
[Illustration: FIG. 230.--AKENES OF BUTTERCUP, one in longitudinal
section.]
A 1-loculed pericarp which dehisces along the front edge (that is, the
inner edge, next the center of the flower) is a =follicle=. The fruit
of the larkspur (Fig. 231) is a follicle. There are usually five of
these fruits (sometimes three or four) in each larkspur flower, each
pistil ripening into a follicle. If these pistils were united, a single
compound pistil would be formed. Columbine, peony, ninebark, milkweed,
also have follicles.
[Illustration: FIG. 231.--FOLLICLE OF LARKSPUR.]
[Illustration: FIG. 233.--CAPSULE OF CASTOR-OIL BEAN AFTER DEHISCENCE.]
[Illustration: FIG. 232.--A BEAN POD.]
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