When the mycelium is about two days old, it begins to form the
long fruiting stalks which we first noticed. To study them, use a
compound microscope magnifying about two hundred diameters. One of
the stalks, magnified, is shown in _a_, Fig. 274. It consists of a
rounded head, the _sporangium, sp_, supported on a long, delicate
stalk, the _sporangiophore_. The stalk is separated from the
sporangium by a wall which is formed at the base of the sporangium.
This wall, however, does not extend straight across the thread, but
it arches up into the sporangium like an inverted pear. It is known
as the _columella, c_. When the sporangium is placed in water, the
wall immediately dissolves and allows hundreds of spores, which
were formed in the cavity within the sporangium, to escape, _b_.
All that is left of the fruit is the stalk, with the pear-shaped
columella at its summit, _c_. The spores that have been set free by
the breaking of the sporangium wall are now scattered by the wind
and other agents. Those that lodge in favorable places begin to grow
immediately and reproduce the fungus. The others soon perish.
[Illustration: FIG. 273.--MUCOR, showing formation of zygospore on the
right; germinating zygospore on the left.]
[Illustration: FIG. 274.--MUCOR.
_a_, sporangium; _b_, sporangium bursting; _c_, columella.]
The mucor may continue to reproduce itself in this way indefinitely,
but these spores are very delicate and usually die if they do not
fall on favorable ground, so that the fungus is provided with another
means of carrying itself over unfavorable seasons, as winter. This
is accomplished by means of curious _thick-walled resting-spores_
or _zygospores_. The zygospores are formed on the mycelium buried
within the substance on which the plant grows. They originate in the
following way: Two threads that lie near together send out short
branches, which grow toward each other and finally meet (Fig. 273).
The walls at the ends, _a_, then disappear, allowing the contents to
flow together. At the same time, however, two other walls are formed
at points farther back, _b_, _b_, separating the short section, _c_,
from the remainder of the thread. This section now increases in size
and becomes covered with a thick, dark brown wall ornamented with
thickened tubercles. The zygospore is now mature and, after a period
of rest, it germinates, either producing a sporangium directly or
growing out as mycelium.
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