=Movements: the Water-tube System.=--The water, which is filtered
through the perforated madreporite, is needed to supply a _system of
canals_ (Fig. 56). The madreporite opens into a canal called the _stone
canal_, the wall of which is hardened by the same kind of material
as that found in the skin. The stone canal leads to the ring canal
which surrounds the mouth (Fig. 56). The ring canal sends _radial
canals_ into each ray to supply the double row of _tube feet_ found in
the groove at the lower side of each ray (Fig. 57). Because of their
arrangement in rows, the feet are also called _ambulacral_ feet (Latin
_ambulacra_, “forest walks”). There is a water holder (_ampulla_), or
muscular water bulb at the base of each tube foot (Fig. 58). These
contract and force the water into the tube feet and extend them. The
cuplike ends of the tubes cling to the ground by suction. The feet
contain delicate muscles by which they contract and shorten. Thus the
animal pulls itself slowly along, hundreds of feet acting together.
The tube feet, for their own protection, may contract and retire into
the groove, the water which extended them being sent back into the
ampulla. This system of water vessels (or water-vascular system) of the
echinodermata is characteristic of them; _i.e._ is not found elsewhere
in the animal kingdom. The grooves and the plates on each side of them
occupy the _ambulacral areas_. The rows of spines on each side of the
grooves are freely movable. (What advantage?) The spines on the aboral
surface are not freely movable.
[Illustration: FIG. 57.--Starfish, from below; tube feet extended.]
[Illustration: FIG. 58.--SECTION OF ONE RAY and central portion of
starfish.
_f1_, _f2_, _f3_, tube feet more or less extended; _au_, eye spot;
_k_, gills; _da_, stomach; _m_, madreporite; _st_, stone canal; _p_,
ampulla; _ei_, ovary.]
=Respiration.=--The _system of water vessels serves the additional
purpose_ of bringing water containing oxygen into contact with various
parts of the body, and the starfish was formerly thought to have no
special respiratory organs. However there are holes in the aboral wall
through which the folds of the delicate lining membrane protrude. These
are now supposed to be _gills_ (_k_, Fig. 58).
=The nervous system= is so close to the aboral surface that much of
it is visible without dissection. Its chief parts are a _nerve ring_
around the mouth, which sends off a _branch_ along each ray. These
branches may be seen by separating the rows of tube feet. They end in a
pigmented cell at the end of each ray called the _eye-spot_.
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