The shells were held together by two large _adductor muscles_. The
anterior adductor (Fig. 193) is near the front end, above the foot.
The posterior adductor is toward the rear end, but not so near the end
as the anterior. Can you find both _muscle scars_ in the shells? Are
they nearer the ventral or dorsal surface? The points of attachment
traveled downward and farther apart as the animal grew (see Fig. 191).
Higher than the larger scars are small scars, or impressions, where the
protractor and retractor muscles that extend and draw in the foot were
attached.
[Illustration: FIG. 193.--ANATOMY OF MUSSEL. (Beddard.)]
The muscular _foot_ extends downward in the middle, halfway between
the shells (Fig. 193). On each side of the foot and behind it hang
down the two pairs of =gills=, the outer pair and the inner pair (Fig.
192). They may be compared to four V-shaped troughs with their sides
full of holes. The water enters the troughs through the holes and
overflows above. Is there a marked difference in the size of the two
pairs of gills? A kind of chamber for the gills is made by the joining
of the mantle flaps below, along the ventral line. The mantle edges
are separated at two places, leaving openings called _exhalent_ and
_inhalent siphons_.
Fresh water with its oxygen, propelled by _cilia_ at the opening and on
the gills, enters through the lower or inhalent siphon, passes between
the gills, and goes to an upper passage, leaving the gill chamber by
a slit which separates the gills from the foot. For this passage, see
arrow (Fig. 194). The movement of the water is opposite to the way the
arrow points. After going upward and backward, the water emerges by the
exhalent siphon. The gills originally consisted of a great number of
filaments. These are now united, but not completely so, and the gills
still have a perforated or lattice structure. Thus they present a large
surface for absorbing oxygen from the water.
[Illustration: FIG. 194.--MUSSEL.
_A_, left shell and mantle flap removed.
_B_, section through body.
=Question:= Guided by other figures, identify the parts to which lines
are drawn.]
The =mouth= is in front of the foot, between it and the anterior
adductor muscle (Fig. 194). On each side of the mouth are the _labial
palps_, which are lateral lips (Fig. 195). They have cilia which convey
the food to the mouth after the inhalent siphon has sent food beyond
the gill chamber and near to the mouth. Thus both food and oxygen enter
at the inhalent siphon. The foot is in the position of a lower lip, and
if regarded as a greatly extended lower lip, the animal may be said to
have what is to us the absurd habit of using its lower lip as a foot.
The foot is sometimes said to be hatchet-shaped (Fig. 195). Do you see
any resemblance? Does the foot penetrate deep or shallow into the sand?
(Fig. 190.) Why, or why not?
[Illustration: FIG. 195.--MUSSEL. From below. Level cut across both
shells.
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