Evolution; Knowledge, Theory of; Philosophy and religion
Let us turn now to the evidence furnished by organic bodies. In place of
deductive illustrations like the foregoing, we shall here find numerous
illustrations which, as being inductively established, are less open to
criticism. The process of mammalian development, for example, will
supply us with numerous proofs ready-described by embryologists.
The first change which the ovum of a mammal undergoes, after
continued segmentation has reduced its yelk to a mulberry-like mass, is
the appearance of a greater definiteness in the peripheral cells of this
mass: each of which acquires a distinct enveloping membrane. These
peripheral cells, vaguely distinguished from the internal ones both by
their greater completeness and by their minuter subdivision, coalesce to
form the blastoderm or germinal membrane. One portion of the blastoderm
presently becomes contrasted with the rest, through the accumulation of
cells still more subdivided, which, together, form an opaque roundish
spot. This _area germinativa_, as it is called, is not sharply
delineated, but shades off gradually into the surrounding parts of the
blastoderm; and the _area pellucida_, subsequently formed in the midst
of this germinal area, is similarly without any precise margin. The
“primitive trace,” which makes its appearance in the centre of the _area
pellucida_, and is the rudiment of that vertebrate axis which is to be
the fundamental characteristic of the mature animal, is shown by its
name to be at first indefinite—a mere trace. Beginning as a shallow
groove, this becomes slowly more pronounced: its sides grow higher,
their summits overlap, and at last unite; and so the indefinite groove
passes into a definite tube, forming the vertebral canal. In this
vertebral canal the leading divisions of the brain are at first
discernible only as slight bulgings; while the vertebræ commence as
indistinct modifications of the tissue bounding the canal.
Simultaneously, the outer portion of the blastoderm has been undergoing
separation from the inner portion: there has been a division into the
serous and mucous layers—a division at the outset indistinct, and
traceable only about the germinal area, but which insensibly spreads
throughout nearly the whole germinal membrane, and becomes definite.
From the mucous layer, the development of the alimentary canal proceeds
as that of the vertebral canal does from the serous layer. Originally a
simple channel along the under surface of the embryonic mass, the
intestine is rendered step by step more distinct by the bending down, on
each side, of ridges which finally join to form a tube—the permanent
absorbing surface is by degrees clearly cut off from that temporary
absorbing surface of which it was at first a part like all the rest. And
in an analogous manner the entire embryo, which at first lies outspread
upon the surface of the yelk-sack, gradually rises up from it, and, by
the infolding of its ventral surface, becomes a separate mass, connected
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