Form and Function: A Contribution to the History of Animal MorphologyRussell, E. S. (Edward Stuart)
History
Form and Function: A Contribution to the History of Animal Morphology
Russell, E. S. (Edward Stuart)
Anatomy, Comparative -- History; Morphology (Animals) -- History
Schwann's next step is to discover what are the essential forces active
in the cell, and here he enters the realm of hypothesis. He finds they
can be reduced to two--an attractive force and a metabolic force. The
attractive force is seen in the process of cell-formation, where first
of all the nucleolus is formed by a concentration and precipitation of
substances found free in the cytoblastem, and in the same way the
nucleus and later the cell are laid down as concentric precipitates from
the cytoblastem. Cell-formation also involves the second or metabolic
force, by means of which the cell alters the chemical composition of the
medium surrounding it so as to prepare it for assimilation. Schwann's
attractive force brings about the actual taking up of the prepared
substance; his metabolic force is the cause of the digestion of food
substances, and is nearly identical with enzyme action. With what
inorganic process, he now asks (p. 239), can the process of
cell-formation be most nearly compared, and the answer obviously is,
with the process of crystallisation. Cells are, it is true, quite
different in shape and consistency from crystals, and they grow by
intussusception, not by apposition--their plastic or attractive forces
seem therefore to be different. A still more important difference is
that the metabolic force is peculiar to the cell. Yet there are
important analogies between crystals and cells. They agree in the
important respect that they both grow in solutions at the cost of the
dissolved substance, according to definite laws, and develop a definite
and characteristic shape. It might even be maintained, Schwann thinks,
that the attractive force of crystals is really identical with that of
cells, and that the difference in result is due merely to the difference
between the substance of the cell and the substance of the crystal. He
points out how organic bodies are remarkable for their powers of
imbibition, and he seeks to show that the cell is the form under which a
body capable of imbibition must necessarily crystallise, and that the
organism is an aggregate of such imbibition-crystals. The analogy
between crystallisation and cell-formation he works out in the following
manner:--"The substance of which cells are composed possesses the power
of chemically transforming the substance with which it is in immediate
contact, in somewhat the same way as the well-known preparation of
platinum changes alcohol into acetic acid. Each part of the cell
possesses this property. If now the cytoblastem is altered by an already
formed cell in such a way that a substance is formed that cannot become
part of the cell, it crystallises out first as the nucleolus of a new
cell. This in its turn alters the composition of the cytoblastem. A part
of the transfomed substance may remain in solution in the cytoblastem or
may crystallise out as the beginning of a new cell; another part, the
cell-substance, crystallises round the nucleolus. The cell-substance is
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