Form and Function: A Contribution to the History of Animal MorphologyRussell, E. S. (Edward Stuart)
History
Form and Function: A Contribution to the History of Animal Morphology
Russell, E. S. (Edward Stuart)
Anatomy, Comparative -- History; Morphology (Animals) -- History
Milne-Edwards frankly takes up the teleological standpoint, and
interprets organic forms on the assumption that they are purposive and
rationally constructed. "To arrive at an understanding of the harmony of
the organic creation," he writes, "it seemed to me that it would be well
to accept the hypothesis that Nature has gone about her work as we would
do ourselves according to the light of our own intelligence, if it were
given us to produce a similar result. Comparing and studying living
things as if they were machines created by the industry of man, I have
tried to grasp the manner in which they might have been invented, and
the principles whose application would have led to the production of
such an assemblage of diversified instruments" (p. 435). The problem is
to discover the laws which rule the diversity of organic forms. The
first and most obvious of these laws is the "law of economy," or the law
of unity of type. Nature, as Cuvier pointed out, has not had recourse to
all the possible forms and combinations of organs; she appears to work
with a limited number of types and to get the greatest possible
diversity out of these by varying the proportions of the constitutive
materials of structure. Within the limits of each type Nature has
brought about diversity by raising her creatures to different degrees of
perfection. This is the second law of organic form, and it is this law
that Milne-Edwards chiefly elaborates. Degrees of perfection mean for
him, as for Aristotle, primarily degrees of perfection of function, but
since structure is necessarily in close relation with function,
perfection of function brings in its train increased perfection of
organisation. This can only be attained by a division of labour[306] among
the organs and by their consequent differentiation. An animal is like a
workshop where some complicated product is manufactured, and the organs
are like the workmen. Each workman has his own special piece of work to
do, at which he becomes thoroughly expert; and the finished product is
manufactured more rapidly and efficiently by the co-operation of workers
each skilled in one department than it would be if each workman had to
produce the whole. Applied to the organism this principle of the
division of labour means the differentiating out of the separate
functions, their localisation in different parts of the organism, and
their co-ordination to produce a combined result.
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