Form and Function: A Contribution to the History of Animal MorphologyRussell, E. S. (Edward Stuart)
History
Form and Function: A Contribution to the History of Animal Morphology
Russell, E. S. (Edward Stuart)
Anatomy, Comparative -- History; Morphology (Animals) -- History
The fundamental forms of living things are, however, merely so many
themes on which a multitude of further variations are woven, through the
action of the laws which rule the detail of organic diversities. These
further laws may be set down under four main heads. Under the first
comes the law of the existence of certain fundamentally distinct
structural types, which are distinguished from one another by their
ground-form, by the number of organ-systems, and by the number of
homotypic organs they possess, but principally by the relative position
of the organs to one another (principle of connections). The form and
connections of the nervous system are of particular importance in
distinguishing the types (_cf._ Cuvier). The second factor in the
diversity of organic form is the action of certain laws of progressive
development[313] (_Entwickelungsgesetze_), which bear the same relation to
the development of the animal kingdom as the laws of individual
development bear to the development of the embryo, for organs appear in
the different animal series in much the same order and manner as they
develop in the individual. These laws are (1) progressive
differentiation of functions and organs; (2) numerical reduction of
serially repeated parts; (3) concentration of functions and their organs
in particular parts of the body; (4) centralisation of organ-systems and
parts of such, so that they come to depend upon one central organ; (5)
internalisation of the "noblest" organs, unless these are necessarily
external, and (6) increase in size of the whole or of parts. Of these
the law of differentiation is by far the most important, and most of the
others are in a sense merely special cases of this fundamental law. To
this law of differentiation is due the increase in complexity or
perfection of organisation which is shown by all the animal series.
Bronn himself recognised the great similarity of this law of progressive
differentiation to Milne-Edwards' principle of the division of labour;
he seems, however, to have arrived at it independently.
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