Form and Function: A Contribution to the History of Animal MorphologyRussell, E. S. (Edward Stuart)
History
Form and Function: A Contribution to the History of Animal Morphology
Russell, E. S. (Edward Stuart)
Anatomy, Comparative -- History; Morphology (Animals) -- History
The _via media_ adopted by Roux is the analysis of development, not
directly into simple physico-chemical processes, but into more complex
organic processes dependent upon the fundamental properties of living
matter. The aim of _Entwicklungsmechanik_ is defined by Roux to be the
reduction of developmental events to the fewest and simplest
_Wirkungsweisen_, or causal processes.[483] Two classes of causal
processes may be distinguished, as "complex components" and "simple
components" of development. The latter are directly explicable by the
laws of physics and chemistry; the former, while in essence
physico-chemical, are yet so very complicated that they cannot at
present be reduced to physico-chemical terms. The ultimate aim of
_Entwicklungsmechanik_ is to reduce development to its "simple
components," but its main task at the present day and for many years to
come is the analysis of development into its "complex components."
These complex components must be accepted as having much of the validity
of physical and chemical laws. They are mysterious in the sense that
they cannot yet be explained mechanistically, but they are constant in
their action, and under the same conditions produce always the same
effect--hence they may be made the subject of strictly scientific study.
They represent biological generalisations, in their way of equal
validity with the generalisations of physics and chemistry.
The principal "complex components" which Roux recognises are somewhat as
follows:--First come the elementary cell-functions of assimilation and
dissimilation, growth, reproduction and heredity, movement and
self-division (as a special co-ordination of cell-movements). Then at a
somewhat higher level, self-differentiation, and the trophic reaction to
functional stimuli. Components of even greater complexity may also be
distinguished, as, for instance, the biogenetic law. The various
tropisms exhibited in development may be regarded as "directive" complex
components. There must be added, not as being itself a component, but
rather as a mode or peculiar property of all functioning, the
omnipresent faculty of self-regulation.
It will be noticed that Roux's "complex components" are simply the
general properties or functions of organised matter.
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