Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In 1878 Munier-Chalmas expressed the opinion, which was supported by
strong evidence, that _Ovulites_ should be referred to the siphoneous
algae[290]. He regarded it as generically identical with _Penicillus_
(_Coralliodendron_, Kützing). It has already been pointed out that
in _Penicillus_ the apical tuft of filaments is partially calcareous
(fig. 33, O)[291]. The individual calcareous segments agree almost
exactly with the fossil _Ovulites_. As a rule the Ovulites occur
as separate egg- or rod-like bodies, but Munier-Chalmas informs me
that occasionally two or three have been found joined end to end in
their natural position. The terminal holes in the fossil specimens
represent the apertures left after the detachment of the calcareous
segments from the uncalcified filaments of the alga. The segments
with two holes at the broader end were no doubt situated at the base
of dichotomising branches as shown in fig. 33, K. The restoration of
_Ovulites_, shown in fig. 33, K, bears a striking resemblance to the
figure of an Australian _Penicillus_ given by Harvey in his _Phycologia
Australica_[292].
It is probable that these Eocene forms agreed closely in habit with
the recent species of _Penicillus_. The portions preserved as fossils
are segments of the filaments which probably formed a terminal brush
of fine branches supported on a stem. The retention of the original
generic name _Ovulites_ is on the whole better than the inclusion of
the fossil species in the recent genus. The Tertiary species lived in
warm seas of the Lower and Middle Eocene of England, Belgium, France
and Italy.
_Halimeda._
An example of an Eocene species of _Halimeda_ has been recorded by
Fuchs from Greifenstein under the name of _Halimeda Saportae_[293]. The
impression has the form of a branched plant consisting of wedge-shaped
or oval segments, and there is a close resemblance to the thallus of a
recent _Halimeda_, e.g. _H. gracilis_ Harv. It is not improbable that
Fuchs’ determination is correct, but without more definite evidence
than is afforded by a mere impression it is a little rash to make use
of the recent generic name.
γ. Dasycladaceae.
In this family of Siphoneae are included a number of genera represented
by species living in tropical and subtropical seas.
The thallus consists of an elongated axial cell fixed to the substratum
by basal rhizoids, and bearing whorls of lateral appendages of limited
growth which may be either simple or branched. Many of the lateral
branches bear sporangia or spores. The thallus is in many species
encrusted with carbonate of lime.
The two genera _Acetabularia_ and _Cymopolia_ may be briefly described
as recent types which are represented by trustworthy fossil forms.
[Illustration: FIG. 34. _Acetabularia mediterranea_ Lamx. From a
specimen in the Cambridge Botanical Museum (nat. size).]
_Acetabularia._ Figs. 33, I, and 34.
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