Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
_Solenopora compacta_ (Billings). Fig. 38. This species was originally
described by Billings as _Stromatopora compacta_, and afterwards
defined by Nicholson and Etheridge. The thallus forms sub-spheroidal
masses, from the size of a hemp-seed to that of an orange. The external
surface is lobulate; the fractured surface has a porcellaneous and
sometimes a fibrous appearance, and is usually white or light brown
in colour. In vertical section (fig. 38, B) the cells are elongated
and arranged in a radiating and parallel fashion; they often occur in
concentric layers. The cells have a diameter of about ¹⁄₁₇ mm. and
possess distinctly undulating walls, as seen in a tangential section
(fig. 38, A). Brown describes certain larger cells in the thallus (fig.
38, A) as sporangia[363], but it is difficult to recognise any distinct
sporangial cavities in the drawing. The example figured is from the
Trenton limestone of Canada; a variety of the same species has been
recorded from the Ordovician rocks of Girvan in Ayrshire. There appear
to be good reasons for accepting Brown’s conclusion that _Solenopora_
belongs to the Corallinaceae rather than to the Hydrozoa, among which
it was originally included. After comparing _Solenopora_ with recent
genera of Florideae, Brown concludes that “the forms of the cells and
cell-walls, the method of increase, and the arrangement of the tissue
cells in the various species of _Solenopora_ bear strong evidence of
relationship between that genus and the calcareous algae[364].”
[Illustration: FIG. 38. Solenopora compacta (Billings). A. Tangential
section. × 100. B. Vertical section. × 50. (After Brown.)]
The importance of the calcareous Rhodophyceae has been frequently
emphasised by recent researches, and our knowledge of the rock-building
forms is already fairly extensive. We possess evidence of the existence
of species of different genera in Ordovician seas, as well as in those
of the Silurian, Triassic, Jurassic, and more recent periods. It is
reasonable to prophesy that further researches into the structure
of ancient limestones will considerably extend our knowledge of the
geological and botanical history of the Corallinaceae.
Numerous fossils have been described as examples of other genera[365]
of Rhodophyceae than those included in the Corallinaceae, but these
possess little or no scientific value and need not be considered.
E. PHAEOPHYCEAE (BROWN ALGAE).
Olive-brown algae, thallus often leathery in texture, composed of
cell-filaments or parenchymatous tissue, in some cases exhibiting
a considerable degree of internal differentiation. The sexual
reproductive organs may be either in the form of passive egg-cells
and motile antherozoids or of motile cells showing no external sexual
difference.
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