Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The list of fossil fungi given by Meschinelli in Saccardo’s _Sylloge
Fungorum_[411] includes certain species which are of no botanical
value, and should have no place in any list which claims to be
authentic.
Among the numerous examples of fossil ‘fungi’ which have no claim to
be classed with plants, there are some which are in all probability
the galleries of wood-eating animals. The radiating grooves frequently
found on the inner face of the bark of a pine tree made by species of
the beetle _Bostrychus_ might be mistaken for the impressions of the
firm strands of mycelial tissue of some Basidiomycetous fungus.
In some notes on fossil fungi by J. F. James[412] contributed to
the American Journal of Mycology in 1893, it is pointed out that a
supposed fungus described by Lesquereux from the Lower Coal-Measures
as _Rhizomorpha Sigillariae_[413], bears a strong likeness to some
insect-burrows, such as those of _Bostrychus_.
“A new fungus from the Coal-Measures” described by Herzer in 1893[414]
may probably be referred to animal agency. In any case there is no
evidence as to the fungoid nature of the object represented in the
figure accompanying Herzer’s description.
[Sidenote: BASIDIOMYCETES.]
More trustworthy evidence of fossil fungi is afforded by the marks
of disease in petrified tissue and by the presence of true mycelia.
In examining closely the calcareous and siliceous plant-tissues from
the Coal-Measures and other geological horizons, one occasionally
sees fine thread-like hyphae ramifying through the cells or tracheal
cavities; in many cases the hyphae bear no reproductive organs and
cannot as a rule be referred to a particular type of fungus. If the
hyphal filaments are unseptate, they most likely belong to some
Phycomycetous species; or if they are obviously septate the Mesomycetes
or the Mycomycetes are the more probable groups. Occasionally there
may be found indications of the characteristic _clamp-connections_ in
the septate filaments; a small semicircular branch, which is given off
from a mycelium immediately above a transverse wall, bends round to
fuse with the filament just below the septum, thus serving as a small
loop-line connecting the cell-cavity above and below a cross wall. Such
clamp-connections are usually confined to the hyphae of Basidiomycetes
and thus serve as a useful aid in identification. A good example of a
clamp-connection in a fossil mycelium is figured by Conwentz[415] in
his monograph on the Baltic amber-trees of Oligocene age. The stout
and thick type of hypha found in some fossil woods agrees closely with
that of _Polyporus_, _Agaricus melleus_ and other well-known recent
Basidiomycetes.
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