Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The generic name was proposed by Vaillant in 1719[438], and adopted by
Linnaeus, who classed the Stoneworts with aquatic phanerogams. As long
ago as 1623[439] a figure of _Chara_ was published by Caspar Bauhin
as a form of _Equisetum_. The generic name _Chara_ has usually been
applied to recent and fossil species alike. The existing species have
a wide distribution; _Chara foetida_, A. Br., a common British form,
occurs in practically all parts of the world. Stems and calcareous
‘fruit-cases’ occur fairly commonly in a fossil state, and differ but
little from recent species, at least as regards essential features.
It is difficult to say at what geological horizon the Stoneworts
are first represented. The first certain traces of _Chara_ occur in
Jurassic rocks, but certain spirally marked subspherical bodies have
been recorded from Devonian and Carboniferous strata, which closely
resemble Chara oogonia, and may be Palaeozoic representatives of the
genus.
In 1889 Mr Knowlton[440] of the American Geological Survey described
some ‘problematic organisms’ found in Devonian rocks at the falls of
the Ohio. Examples of these fossils are shown in fig. 46 _b_ and _c_;
the spirally grooved body measures from 1·50 to 1·80 mm. in diameter,
and about 1·70 mm. in length. The Chara-like character of the fossils
had been previously suggested by Meek[441] in 1873. Without going into
the arguments for or against placing these fossils in the Chareae,
they may at least be mentioned as possible but not certain Palaeozoic
forms of _Chara_ or an allied genus.
[Illustration: FIG. 46. _a_. _Chara Bleicheri_ Sap. × 30. _b_ and _c_.
Devonian _Chara_? sp. _circa_ × 12. _d_ and _e_. _Chara Wrighti_
Forbes. _circa_ × 12.]
1. _Chara Bleicheri_, Saporta. Fig. 46, _a_.
In this form the ‘fruits’ are minute and subspherical, ·39-·44 mm.
long, and ·35-·40 mm. broad, showing in side view 5–6 slightly oblique
spiral bands. Each spiral band bears a row of slightly projecting
tubercles.
This species was first described by Saporta[442] from the Oxfordian
(Jurassic) rocks of the Department of Lot in France; it is compared by
the author of the species with _Chara Jaccardi_ Heer, described by Heer
from the Upper Jurassic rocks of Switzerland.
2. _C. Knowltoni_, Seward. Fig. 45, _a_ and _b_, and Fig. 47.
The Oogonia are broadly oval, about ·5mm. in length, and at the
broadest part of about the same breadth. The surface is marked by
eleven or twelve bands in the form of a flattened spiral. The stems
possess investing cortical cells.
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