Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The small drawing in fig. 55, 3 shows part of the ring of thick
nodal wood; the section cuts through two bundles about their point
of bifurcation, the strand _x_ is passing out in a radial direction
to a lateral branch, the strand to the right of _x_ and the separate
fragment of a strand to the left of _x_ are portions of leaf-trace
bundles on their way to the leaf-sheath. Reverting to fig. 54, B, the
other structures seen in the section are the leaf-sheaths (_l_ and
_m_), the vallecular canal (_f_), the epidermis, cortex and pith (_k_,
_e_ and _a_) of the stem. The epidermis which has been ruptured by the
root and branch is indicated at _i_, _i_; the dotted lines traversing
the upper part of the pith of the lateral branch mark the position of a
nodal diaphragm.
[Illustration: FIG. 55. 1. Transverse section of a root of _Equisetum
variegatum_ Schl., _e_ endodermis, or outer layer of the
phloeoterma (after Pfitzer; × 160). 2. Transverse section of
rhizome of _E. maximum_, slightly enlarged. 3. Transverse section
through a node of _E. maximum_, _x_, branch of vascular strand
(slightly enlarged). 4. Transverse section through a node of _E.
maximum_ showing the mass of xylem, _px_ protoxylem (× 175). (Figs.
3 and 4 after Cormack.)]
Immediately external to each vascular strand, as seen in transverse
section, there is a layer of cells containing starch, and this
is followed by a distinct endodermis, of which the cells show the
characteristic black dot in the cuticularised radial walls (fig. 52,
D). Beyond the endodermis there is the large-celled parenchyma of
the rest of the cortex. Tannin cells occur here and there scattered
among the ground tissue. On the same radius on which each vascular
strand occurs, the cortical parenchyma passes into a mass of
sub-epidermal thick-walled mechanical tissue or stereome. Alternating
with the ridges of stereome, the grooves are occupied by thin-walled
chlorophyll-containing tissue which carries on most of the assimilating
functions, and communicates with the external atmosphere by means of
stomata arranged in vertical rows down each internode. The continuity
of the cortical tissue is interrupted by the occurrence of large
longitudinal _vallecular canals_ alternating in position with the stem
ridges and vascular strands (fig. 52, C, _v_). The epidermis consists
of a single layer of cells, containing stomata, and with the outer
cell-walls impregnated with silica.
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