Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In dealing with the fossil Equisetales, we will first consider
the genera _Equisetites_, _Phyllotheca_ and _Schizoneura_, and
afterwards describe the older and better known genera _Calamites_
and _Archaeocalamites_. A thoroughly satisfactory classification of
the members of the Equisetales is practically impossible without
more data than we at present possess. It has been the custom to
include _Equisetites_, _Phyllotheca_ and _Schizoneura_ in the family
Equisetaceae, and to refer _Calamites_ and _Archaeocalamites_ to the
Calamarieae; such a division rests in part on assumption, and cannot
be considered final. When we attempt to define the Equisetales and the
two families Equisetaceae and Calamarieae, we find ourselves seriously
hampered by lack of knowledge of certain important characters, which
should be taken into account in framing diagnoses. There is little
harm in retaining provisionally the two families already referred to,
if we do not allow a purely arbitrary classification to prejudice our
opinions as to the affinities of the several members of the Equisetales.
The Equisetaceae might be defined as a family including plants which
were usually herbaceous but in some cases arborescent, bearing
verticils of leaves in the form of sheaths more or less deeply divided
into segments or teeth. The strobili were isosporous and consisted of
a central axis bearing verticils of distally expanded sporophylls with
sporangia, as in _Equisetum_. The genus _Equisetites_ might be included
in this family, but it must be admitted that we know next to nothing
as to its anatomy, and we cannot be sure that the strobili were always
isosporous.
The genus _Schizoneura_ is too imperfectly known to be defined with any
approach to completeness, or to be assigned to a family defined within
certain prescribed limits. _Phyllotheca_ is another genus about which
we possess but little satisfactory knowledge; we are still without
evidence as to its structure, and the descriptions of the few strobili
that are known are not consistent. Recent work points to a probability
of _Phyllotheca_ being closely allied to _Annularia_, a genus included
in the Calamarieae, and standing for a certain type of Calamitean
foliage-shoots.
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