Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Mr Kidston[503] founded this species on a few specimens of cones
found in the Middle Coal-Measures of Barnsley in Yorkshire. The best
example of the cone described by Kidston has a length of 2·5 cm.,
and a breadth of 1·5 cm.; the surface is divided up into several
hexagonal areas 4 mm. high and 5 mm. wide. Each of these plates shows
a fairly prominent projecting point in its centre; this is regarded
as the point of attachment of the sporangiophore axis which expanded
distally into a hexagonal plate bearing sporangia. An examination of
Mr Kidston’s specimens enabled me to recognise the close resemblance
which he insists on between the fossils and such a recent Equisetaceous
strobilus as that of _Equisetum limosum_ Sm. Nothing is known of the
structure of the fossils beyond the character of the superficial
pattern of the impressions, and it is impossible to speak with absolute
confidence as to their nature. The author of the species makes
use of the generic name _Equisetum_; but in view of our ignorance
of structural features it is better to adopt the more usual term
_Equisetites_.
Since Kidston’s description was published I noticed a specimen in the
British Museum collection which throws some further light on this
doubtful fossil. Part of this specimen is shown in fig. 57, _A_. The
stem is 21 cm. in length and about 5 mm. broad; it is divided into
distinct nodes and internodes; the former being a little exaggerated in
the drawing. The surface is marked by fine and irregular striations,
and in one or two places there occur broken pieces of narrow linear
leaves in the neighbourhood of a node. Portions of four cones occurring
in contact with the stem, appear to be sessile on the nodes, but
the preservation is not sufficiently good to enable one to speak
with certainty as to the manner of attachment. Each cone consists of
regular hexagonal depressions, which agree exactly with the surface
characters of Kidston’s type-specimen. The manner of occurrence of the
cones points to a lateral and not a terminal attachment. The stem does
not show any traces of Equisetaceous leaf-sheaths at the nodes, and
such fragments of leaves as occur appear to have the form of separate
linear segments; they are not such as are met with on _Equisetites_.
It agrees with some of the slender foliage-shoots of Calamitean plants
often described under the generic name _Asterophyllites_. As regards
the cones; they differ from the known Calamitean strobili in the
absence of sterile bracts, and appear to consist entirely of distally
expanded sporophylls as in _Equisetum_. The general impression afforded
by the fossil is that we have not sufficient evidence for definitely
associating this stem and cones with a true _Equisetites_. We may,
however, adhere to this generic title until more satisfactory data are
available.
2. _Equisetites spatulatus_ Zeill. Fig. 58, _A_.
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