Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 1 : $b [A text-book] for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
There are a few fossil stems from Permian rocks of Siberia,
from Jurassic strata in Italy, and from Lower Mesozoic and
Permo-Carboniferous beds in South America, South Africa, India and
Australia which do not conform in all points to the usually accepted
definition of _Equisetites_, and so justify their inclusion in an
allied genus. On the other hand there are numerous instances of stems
or branches which have been referred to _Phyllotheca_ on insufficient
grounds. Our knowledge of this Equisetaceous plant has recently been
extended by Zeiller[549], who has recorded its occurrence in the
Coal-Measures of Asia Minor associated with typical Upper Carboniferous
plants. The same author[550] has also brought forward good evidence for
the Permian age of the beds in Siberia and Altai, where _Phyllotheca_
has long been known. It is true that Zigno’s species of the genus
occurs in Italian Jurassic rocks, but on the whole it would seem that
this genus is rather a Permian than a Jurassic type. The species
which Zeiller describes under the name _Phyllotheca Rallii_ from
the Coal-Measures of Herakleion (Asia Minor) shows some points of
contact with _Annularia_. It is much to be desired, however, that
we might learn more as to the reproductive organs of this member of
the Equisetales; until we possess a closer acquaintance with the
fructification we cannot hope to arrive at any satisfactory conclusion
as to the exact position of the genus among the Calamarian and
Equisetaceous forms. M. Zeiller[551] informs me that his specimens of
_P. Rallii_, which are to be fully described in a forthcoming work,
include fossil strobili resembling those of _Annularia radiata_.
The verticils of linear leaves fused basally into a sheath agree
in appearance with the star-like leaves of _Annularia_, but in
_Phyllotheca Rallii_ the segments appear to spread in all directions
and are not extended in one plane as in the typical _Annularia_[552].
1. _Phyllotheca deliquescens_ (Göpp.).
In an account of some fossil plants collected by Tchikatcheff in
Altai, Göppert[553] describes and figures two imperfect stems of an
Equisetum-like plant. Owing to the apparent absence of nodal lines on
the surface of the stem the generic name _Anarthrocanna_ is proposed
for the fossils; and the manner in which the main axis appears to break
up into slender branches suggested the specific name _deliquescens_.
Schmalhausen[554] afterwards recognised the generic identity of
Göppert’s fragments with the Indian and Australian stems referred to
the genus _Phyllotheca_ by McCoy[555] and Bunbury[556].
We may define the species as follows:—
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